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Bison (Bison bison) close up head portrait with snow on fur, Yellowstone National Park, USA, February
Chamois (Rupicapra rupicapra), Ballons des Vosges Regional Natural Park, Vosges, Lorraine, France, December
Portrait of male Reindeer (Rangifer tarandus) with large antlers, in velvet. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, June
Pyrenean chamois {Rupicapra rupicapra pyrenaica} Pyrenees, France
Three Llamas (Lama glama) resting in the Machu Picchu ruins. Urubamba, Peru. August, 2016
Red deer (Cervus elaphus) stag bellowing. Surrey, UK, October
Red deer (Cervus elaphus) female amongst autumn foliage, Arran, Scotland, October 2013
American Bison (Bison bison) in habitat, Grand Teton National Park, Wyoming, USA, October
Masai giraffes (Giraffa camelopardalis tippelskirchi), at sunrise, Masai-Mara Game Reserve, Kenya. March
Female Ibex (Capra ibex) portrait, Creux du Van Natural Reserve, Val de Travers, Switzerland, December
Masai giraffes (Giraffa camelopardalis tippelskirchi) in dry season, Masai-Mara Game Reserve, Kenya. March
Wildebeest (Connochaetes taurinus) herd walking in a line in the rain at sunset, Masai-Mara Game Reserve, Kenya. September
Red deer (Cervus elaphus) stag bellowing during rut, Derbyshire, UK
Topi (Damaliscus korrigum / Damaliscus lunatus) territorial male standing on mound looking across savanna. Masai-Mara game reserve, Kenya
Musk ox (Ovibos moschatus) two in habitat, Wrangel Island, Far Eastern Russia, October
Springbok (Antidorcas marsupialis) grazing in front of sand dunes, Namib Naukluft Park, Namibia, June 2006
Musk ox (Ovibos moschatus) portrait of two standing closely, Wrangel Island, Far Eastern Russia, September
Aerial view of two Gemsbok (Oryx gazella) in sand dunes, Namib Desert, Namibia
Chamois (Rupicapra rupicapra) walking with the moon behind, Mercantour National Park, France, October
Alpacas {Lama pacos} Andes. Ecuador
Masai giraffe (Giraffa camelopardalis tippelskirchi) two silhouetted at sunrise, Msai Mara Game Reserve, Kenya
Blue sheep (Pseudois nayaur) Hemis National Park, Jammu and Kashmir, India
Ibex (Capra Ibex) fighting, Aiguilles Rouges National Nature Reserve, Haute-Savoie, France, June
Giraffe (Giraffa camelopardalis) baby, Marakele Private Reserve, Waterberg Biosphere Reserve, Limpopo Province, South Africa
Red deer (Cervus elaphus) stag bellowing, Richmond Park, London, England, UK. October
Saiga antelope (Saiga tatarica) male, standing in grassland in morning sunlight, Betpak Dala, Kazakhstan. Critically endangered
Domestic yak (Bos grunniens) lying under prayer flags near shores of Nam-Tso Lake (at 4, 730m), second largest salt lake in China. Tibet, China
Chamois (Rupicapra rupicapra) head portrait, Schwyz, Switzerland. November
Ibex (Capra ibex) feeding on grass, head portrait, Schwyz, Switzerland. August
Two Ibex (Capra ibex) males, standing on hillside, Lucerne, Switzerland. August
Iberian ibex (Capra pyrenaica), pair with male top right, during rutting season. Sierra Nevada National Park, Andalusia, Spain. October
Iberian ibex (Capra pyrenaica) kid peering over rocks. Sierra Nevada National Park, Andalusia, Spain. June
Iberian ibex (Capra pyrenaica) yearling making its way down snow covered slope. Sierra Nevada National Park, Andalusia, Spain. February
Two Iberian ibex (Capra pyrenaica), adult males, fighting. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), female, being pestered by kid, aged two months, begging for milk. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica) kid, aged two months, making its way down cliff, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), female, grazing, being pestered by kid, aged two months, begging for milk, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July
Four Iberian ibex (Capra pyrenaica) kids, aged two months, playing, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), adult male, sitting down in snow covered landscape during snowfall. Sierra Nevada National Park, Andalusia, Spain. April
Iberian ibex (Capra pyrenaica), alert female with kid aged between one and two months. Sierra Nevada National Park, Andalusia, Spain. May
Iberian ibex (Capra pyrenaica), adult female, walking along coastal cliff. Mediterranean coast, Andalusia, Spain. November
South Andean deer (Hippocamelus bisulcus), adult female with fawn, resting on mountainside. Cerro Castillo National Park, Patagonia, Chile. April
Red deer (Cervus elaphus), young male, swimming through water. Bushy Park, London, UK. September
Masai giraffe (Giraffa camelopardalis tippelskirchi) running across plain. Amboseli National Park, Kenya. July. Digitally enhanced
Wild boar (Sus scrofa) piglet, sniffing the air, La Pampa province, Patagonia, Argentina. Captive
Wild boar (Sus scrofa) piglet nuzzling a dog, La Pampa province, Patagonia, Argentina. Captive
Nilgiri tahr (Nilgiritragus hylocrius) standing next to road signage meant to warn vehicles about the possible road crossings of tahrs in the area, Valparai, Tamil Nadu, India
Springbok (Antidorcas marsupialis) herd in grassland after rain, Palmwag Conservancy, Damaraland, Namibia
