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Gemsbok (Oryx gazella) in sand dunes, Namibia. Finalist in the Mammals Category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
American bison (Bison bison) female covered in hoar frost near hot spring, portrait. Midway Geyser Basin, Yellowstone National Park, USA. February
Herd of American bison (Bison bison) in snow, Yellowstone National Park, Wyoming, Yellowstone, January
Red deer (Cervus elaphus) stag in the snow, Scotland, March
Reindeer (Rangifer tarandus) herd, antlers in velvet, walking across upland moor, Cairngorms, Scotland, UK, August
Red deer (Cervus elaphus) stag in upland landscape. Lochcarron, Highlands, Scotland, UK
RF - Bison (Bison bison) breathing in the cold air, Yellowstone National Park, USA, February
Red deer (Cervus elaphus) backlit at dawn with visible breath. UK. October
Bearded pig (Sus barbatus) close up of snout, Bako National Park, Sarawak, Borneo
Caribou / Reindeer bull (Rangifer tarandus) close-up grazing, Denali National Park, Alaska, USA, September
Moose (Alces alces) bull swimming in water, Baxter State Park, Maine, USA
American bison (Bison bison) Yellowstone National Park, Montana, USA
Red deer (Cervus elaphus) stag silhouetted at sunset, Scotland, UK, February
Siberian musk deer (Moschus moschiferus) male in snow, Irkutsk, Russia. January
Male Saiga antelope (Saiga tatarica) in winter, The Black Lands National Park, Kalmykia, Russia
Alpine ibex (Capra ibex) female, Jura, Switzerland, September
Alpine ibex (Capra ibex), male in habitat, Bernese Alps, Switzerland. July
Guanaco (Lama guanicoe), Punta Norte, Peninsula Valdes, Argentina
RF - Red Deer stag (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Bison (Bison bison) herd walking in snow, Yellowstone National Park, Wyoming, USA, February
Red deer stag (Cervus elaphus) in heavy snow, Cheshire, UK, March
Impala with Oxpeckers. Kruger National Park, South Africa
Gaur (Bos gaurus) Manas National Park UNESCO World Heritage Site, Assam, India
American bison (Bison bison) male walking through deep snow. Hayden Valley. Yellowstone National Park, Wyoming, USA. January
Bearded pig (Sus barbatus) digging in sand, foraging for crabs on beach, Bako National Park, Sarawak, Borneo
Bison (Bison bison) female with calf walking through snow in front of frost-covered forest, Yellowstone National Park, USA
Aerial view of Reindeer (Rangifer tarandus) fording water whilst being herded by Nenet herders. Nenets Autonomous Okrug, Arctic, Russia, July 2017
Asiatic Wild Water Buffalo (Bubalus arnee), Royal Manas National Park, Bhutan
Red deer (Cervus elaphus) male stag roaring portrait, rut season, Jura, Scotland, UK, September
Young Lamas in pasture (Lama glama) altiplano of Atacama Desert, Chile
Red deer stag (Cervus elaphus) during rut on lookout, Exmoor National Park, Somerset / Devon, England. October
Male Nubian Ibex (Capra nubiana) at the Arabian Wildlife Centre captive-breeding project, Sharjah, United Arab Emirates. (captive)
Elk (Cervus canadensis) bull and cw sniffing noses, Yellowstone National Park, Wyoming, USA. September
Rocky mountain elk (Cervus elaphus nelsoni) bull bugling during rut. Rocky Mountain National Park, Colorado, USA. October
Tibetan gazelle / Goa (Procapra picticaudata) Keke Xili / Hoh Xil Nature Reserve, Tibetan High plateau, Qinghai, China
Hippopotamus (Hippopotamus amphibius) with water hyacinth still on back after leaving water, Mana Pools National Park, Zimbabwe
Autumn quaking aspen grove (Populus tremuloides) with trunks scarred by browsing elk (Cervus canadensis). Below Turret Ridge, Uncompahgre National Forest, Colorado, USA. October
White-tailed deer (Odocoileus virginianus) doe standing in snow, New York, USA
Altai argali sheep (Ovis ammon), three grazing on slope. Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China. October
Tibetan gazelle (Procapra picticaudata) in snow. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October
Moose bull (Alces alces) walking in forest clearing, Denali National Park, Alaska, USA, September
Guanacos (Lama guanicoe), two males fighting, trying to over power opponent and bite each others testicles. Torres del Paine National Park, Patagonia, Chile. December
Roe Deer stag (Capreolus capreolus) standing in a woodland glade. Potterick Carr Nature Reserve, Doncaster, Yorkshire, England, UK, July
Silver Dik-dik (Madoqua piacentinii) portrait, Masai Mara, Kenya. March
Guanaco (Lama guanicoe) female, alert whilst looking at Puma (Puma concolor) on on opposite slope, portrait. Torres del Paine National Park, Patagonia, Chile. November
Okapi (Okapia johnstoni). Beauval Zoo Parc, France. Captive
Gayal / Mithun (Bos frontalis) on river bank. Arunachal Pradesh, North East India. November 2014
