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Gemsbok (Oryx gazella) in sand dunes, Namibia. Finalist in the Mammals Category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
American bison (Bison bison) female covered in hoar frost near hot spring, portrait. Midway Geyser Basin, Yellowstone National Park, USA. February
Herd of American bison (Bison bison) in snow, Yellowstone National Park, Wyoming, Yellowstone, January
Red deer (Cervus elaphus) stag in the snow, Scotland, March
Reindeer (Rangifer tarandus) herd, antlers in velvet, walking across upland moor, Cairngorms, Scotland, UK, August
Okapi (Okapia johnstoni). Beauval Zoo Parc, France. Captive
Male Saiga antelope (Saiga tatarica) in winter, The Black Lands National Park, Kalmykia, Russia
Red deer (Cervus elaphus) stag in upland landscape. Lochcarron, Highlands, Scotland, UK
RF - Bison (Bison bison) breathing in the cold air, Yellowstone National Park, USA, February
Red deer (Cervus elaphus) backlit at dawn with visible breath. UK. October
Bearded pig (Sus barbatus) digging in sand, foraging for crabs on beach, Bako National Park, Sarawak, Borneo
Bearded pig (Sus barbatus) close up of snout, Bako National Park, Sarawak, Borneo
Caribou / Reindeer bull (Rangifer tarandus) close-up grazing, Denali National Park, Alaska, USA, September
Moose (Alces alces) bull swimming in water, Baxter State Park, Maine, USA
American bison (Bison bison) Yellowstone National Park, Montana, USA
Red deer (Cervus elaphus) stag silhouetted at sunset, Scotland, UK, February
Siberian musk deer (Moschus moschiferus) male in snow, Irkutsk, Russia. January
Alpine ibex (Capra ibex) female, Jura, Switzerland, September
Alpine ibex (Capra ibex), male in habitat, Bernese Alps, Switzerland. July
Guanaco (Lama guanicoe), Punta Norte, Peninsula Valdes, Argentina
RF - Red Deer stag (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Bison (Bison bison) herd walking in snow, Yellowstone National Park, Wyoming, USA, February
Red deer stag (Cervus elaphus) in heavy snow, Cheshire, UK, March
Impala with Oxpeckers. Kruger National Park, South Africa
American bison (Bison bison) male walking through deep snow. Hayden Valley. Yellowstone National Park, Wyoming, USA. January
Bison (Bison bison) female with calf walking through snow in front of frost-covered forest, Yellowstone National Park, USA
Gaur (Bos gaurus) Manas National Park UNESCO World Heritage Site, Assam, India
Aerial view of Reindeer (Rangifer tarandus) fording water whilst being herded by Nenet herders. Nenets Autonomous Okrug, Arctic, Russia, July 2017
Asiatic Wild Water Buffalo (Bubalus arnee), Royal Manas National Park, Bhutan
Roe Deer stag (Capreolus capreolus) standing in a woodland glade. Potterick Carr Nature Reserve, Doncaster, Yorkshire, England, UK, July
Young Lamas in pasture (Lama glama) altiplano of Atacama Desert, Chile
Red deer stag (Cervus elaphus) during rut on lookout, Exmoor National Park, Somerset / Devon, England. October
Female American buffalo (Bison bison) walking across grassland with newborn calf, Yellowstone National Park, Wyoming, USA. May
Male Nubian Ibex (Capra nubiana) at the Arabian Wildlife Centre captive-breeding project, Sharjah, United Arab Emirates. (captive)
Elk (Cervus canadensis) bull and cw sniffing noses, Yellowstone National Park, Wyoming, USA. September
Rocky mountain elk (Cervus elaphus nelsoni) bull bugling during rut. Rocky Mountain National Park, Colorado, USA. October
Alpine ibex (Capra ibex) resting in mountain pasture, Ticino, Switzerland
Tibetan gazelle / Goa (Procapra picticaudata) Keke Xili / Hoh Xil Nature Reserve, Tibetan High plateau, Qinghai, China
Guanaco (Lama guanicoe) in grassland with Torres del Paine rock towers in background. Torres del Paine National Park, Patagonia, Chile. January 2020
Hippopotamus (Hippopotamus amphibius) with water hyacinth still on back after leaving water, Mana Pools National Park, Zimbabwe
Autumn quaking aspen grove (Populus tremuloides) with trunks scarred by browsing elk (Cervus canadensis). Below Turret Ridge, Uncompahgre National Forest, Colorado, USA. October
White-tailed deer (Odocoileus virginianus) doe standing in snow, New York, USA
Altai argali sheep (Ovis ammon), three grazing on slope. Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China. October
Tibetan gazelle (Procapra picticaudata) in snow. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October
Chital (Axis axis) female leaping. Tadoba Andhari Tiger Reserve / Tadoba National Park, Maharashtra, India
Moose bull (Alces alces) walking in forest clearing, Denali National Park, Alaska, USA, September
Guanacos (Lama guanicoe), two males fighting, trying to over power opponent and bite each others testicles. Torres del Paine National Park, Patagonia, Chile. December
Silver Dik-dik (Madoqua piacentinii) portrait, Masai Mara, Kenya. March
Guanaco (Lama guanicoe) female, alert whilst looking at Puma (Puma concolor) on on opposite slope, portrait. Torres del Paine National Park, Patagonia, Chile. November
Gayal / Mithun (Bos frontalis) on river bank. Arunachal Pradesh, North East India. November 2014
Chinese Water Deer (Hydropotes inermis) Norfolk UK September
Roe deer (Capreolus capreolus) buck with vegetation on antlers in rutting season, Scotland, UK. August
Alpine ibex (Capra ibex) adult males fighting in alpine landscape, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Alpine ibex (Capra ibex) struggling in deep snow on a steep slope, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, January
Snow leopard (Uncia uncia) with Bharal (Pseudois nayaur) prey, Serxu County, Garze Prefecture, Sichuan Province, China
Vicuna (Vicugna vicugna), five running across salt flat with mountain in background. Salar de Ascotan, Chile. September 2018
Red deer (Cervus elaphus) stag, Saxony, Germany. September
Bactrian camels (Camelus bactrianus) on sand, Nubra Valley, Ladakh, India. September
Muskox (Ovibos moschatus), male portrait. Dovrefjell National Park, Norway
Male American Bison (Bison bison) walking through snow. Firehole River Valley, Yellowstone National Park, Wyoming, USA. January
Sable antelope (Hippotragus niger) bull charging through the shallows of the Selinda Spillway, with another behind, northern Botswana, August
Group of four Giraffes (Giraffa camelopardalis) Mkuze, South Africa
Moose (Alces alces) female with twin calves, Baxter State Park, Maine, USA, June
Purebred Berkshire piglet amongst autumn foliage, Smithfield, Rhode Island, USA
Wild boar (Sus scrofa) Vosges, France, July
Warthog (Phacochoerus aethiopicus) female with piglets, Okavango Delta, Botswana
Collared peccary (Tayassu tajacu), adult drinking, South Texas, USA. May
Wild boar (Sus scrofa) eyes glowing at dusk, introduced species in La Pampa, Argentina
Wild boar (Sus scrofa) camera trap image in the Jura mountains, Switzerland, September
Collared peccary (Pecari tajacu) Laredo Borderlands, Texas, USA. April
Warthogs (Phacochoerus aethiopicus) mother with babies drinking, Mkhuze game reserve, KwaZuiu-Natal, South Africa, June
Warthog (Phacochoerus africanus) mother with baby on back sleeping, Masai Mara, Kenya
Wild boar (Sus scrofa) at night with the milky way in the background, Gyulaj, Tolna, Hungary. August. Taken using long exposure with flash at night
Wild boar (Sus scrofa) piglets (known as humbugs ) in woodland clearing. Forest of Dean, Gloucestershire, UK. May
Wild boar (Sus scrofa) in old mixed conifer and broadleaf forest, Punia Forest Reserve, Lithuania, May
RF- Domestic pig (Sus domestica) swimming in sea. Exuma Cays, Bahamas. Tropical West Atlantic Ocean. This family of pigs live on this beach in the Bahamas and enjoy swimming in the sea
White lipped peccary (Tayassu pecari) on forest floor, Lago Preto Conservation Concession, Yavari River, Peru
Springbok (Antidorcas marsupialis) herd fleeing predator, Kgalagadi Transfrontier Park, South Africa
Reindeer (Rangifer tarandus) bull in snow, Cairngorms Reindeer Herd, reintroduced to Scottish Highlands, Cairngorm National Park, Speyside, Scotland, December
Asiatic wild buffalo (Bubalus arnee) portrait of female. Kaziranga National Park, India
Alpine ibex (Capra ibex) close-up portrait. Gran Paradiso National Park, the Alps, Italy. January. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Vicuna (Vicugna vicugna) Uyuni, Altiplano, Bolivia
Llama calf on the altiplano, Bolivia, December 2016
