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Male Leach's sea star (Leiaster leachi) broadcast spawning, releasing streams of sperm from its arms. Kagoshima Prefecture, Kyushu, Japan
Ewens Ponds, spring fed limestone ponds at the Ewens Ponds Conservation Park, South Australia, Australia March 2015. Photographed for The Freshwater Project
Crested newt (Triturus carnifex) Gelderland, The Netherlands, April. Winner of the Amphibians category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
Great grey owl (Strix nebulosa) taking off from snowy ground, Kuusamo, Finland. March
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
Sunset at Trebarwith Strand with incoming tide, North Cornwall, UK. March 2014. *NOT AVAILABLE FOR UK 2016 RETAIL CALENDARS
RF - Reticulated Glass Frog (Hyalinobatrachium valerioi) backlit showing highly translucent body. Osa Peninsula, Costa Rica
Leafless tree canopy silhouetted in winter. Gannochy, Scotland, February
Monochrome image of a Hawthorn tree (Crataegus monogyna) and granite outcrop near Saddle Tor, Dartmoor National Park, Devon, England, UK, January 2011
Star trails above Baobab trees (Adansonia za) at night, Zombitse National Park, Madagascar, June 2010
Ancient dead Camelthorn tree (Vachellia erioloba) trees with red dunes and mist, Namib desert, Deadvlei, Sossusvlei, Namibia. August 2015
Misty Pine (Pinus) forest. Milpa Alta forest, outskirts of Mexico City, Mexico. June
Velvet swimming crab (Necora puber) male, in defensive posture, with wreck of the Rosalie in background, Weybourne, Norfolk, England, UK, North Sea
Crested sculpin (Blepsias bilobus) hiding in smack of Moon jellyfish (Aurelia aurita), Prince William Sound, Alaska, USA, Pacific Ocean
Smack of Moon jellyfish (Aurelia aurita), Prince William Sound, Alaska, USA, Pacific Ocean
Lion's mane jellyfish (Cyanea capillata) preying upon smack of Moon jellyfish (Aurelia aurita), Prince William Sound, Alaska, USA, Pacific Ocean
RF - Schoolmaster snapper (Lutjanus apodus) swimming through underwater caverns within a coral reef, Grand Cayman, Cayman Islands, Caribbean Sea
Dead Langs short-tailed blue butterfly (Leptotes pirithous) in a river with bacterial film, Los Alcornocales Natural Park, Southern Spain, September
Cork trees (Quercus suber) stripped of bark, in forest. Los Alcornocales Natural Park, Cortes de la Frontera, southern Spain, November
Aurora Borealis and coastline at Hornstrandir, Iceland. April 2016
Underwater in a cryoconite hole, a hole formed by windblown dust and particles which warm up ice of glacier and creates meltwater holes
Calcified trees at Tangled Creek in winter, Yellowstone National Park, Wyoming, USA, February 2013
Dead trees in Deadvlei clay pan, Sossusvlei. Namib-Naukluft National Park, Namibia, September 2013
Snowy owl (Bubo scandiaca) female in flight, Canada. February
A long exposure renders Bar jacks (Carangoides ruber) as ghostly trails as they swim around a sponge, on a coral reef, at night. East End, Grand Cayman, Cayman Islands
Herons at night on Lake Csaj, Kiskunsag National Park, Hungary. Winner of the Birds category, Wildlife Photographer of the Year (WPOY) Competition 2014
Aerial view of Amazon Rainforest, and the Manati River, Peru, July 2015
