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Proboscis monkey (Nasalis larvatus) male, Sabah, Borneo
European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition
Eastern lowland gorilla (Gorilla beringei graueri) silverback named Chimanuka, Kahuzi-Biega National Park, South Kivu Province, Democratic Republic of Congo (DRC)
Mountain gorillas (Gorilla beringei) Hirwa group led by the silverback dominant male Munyinya, Volcanoes National Park, Rwanda, elevation 2610 m
Female chimpanzee (Pan troglodytes troglodytes) portrait. Yoko, age 35 years, is the dominant female of the group, Conkouati-Douli National Park, Republic of Congo, Africa
Collie cross dogs Brec and Bliss, exchanging angry snarls. Brec with hackles raised
Lioness (Panthera leo) chasing away Cheetah (Acinonyx jubatus) Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Two Iberian ibex (Capra pyrenaica), adult males, fighting. Sierra Nevada National Park, Andalusia, Spain. July
Two wild Onaqui stallions (Equus ferus) fighting for dominance in between their respective herd. Onaqui Mountain Herd Management Area, Utah, USA. September
Black grouse (Lyrurus tetrix) male, jumping, part of a dominance display, at lek in early morning light, Kuusamo, Finland, April
Two Ruffs (Philomachus pugnax) male, competing at a lek, Vardo, Norway. May
Two male Polar bears (Ursus maritimus) standing on hind legs, sparring, Churchill, Canada. November
Two Red deer (Cervus elaphus) stags fighting, Bradgate Park, Leicestershire, UK. October
Alpha male Grey wolf (Canis lupus) biting angry young wolf to assert dominance, Finland. July
Two male Black grouse (Lyrurus tetrix) displaying at lek, Utajarvi, Finland, April
RF- Gannet (Morus) in flight, Great Saltee, County Wexford, Ireland. April
