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Young Japanese macaque (Macaca fuscata) kissing another macque in the thermal hot springs, Jigokudani, Japan, January
Ground squirrels (Xerus inauris ) one with its tail over the back of another, Kgalagadi Transfrontier Park, Northern Cape, South Africa, February
European Brown Bear (Ursus arctos arctos), a large male taking a rest in a cloud of mosquitoes. Kajaani, Finland. June
Lemon sole (Microstomus kitt) portrait. Gardur, south west Iceland. North Atlantic Ocean
Male Lion (Panthera leo) shaking off water in the rain, Masai-Mara game reserve, Kenya