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Prophet flowers (Arnebia pulchra) with dark spots that act as nectar guides. Caucasus, Russia. June
Chilean monkey flower (Mimulus naiandinus). The lower lip has a groove to guide the pollinators proboscis to the nectar source, with red spots acting as nectar guides
Sacred lotus (Nelumbo nucifera Charles Thomas ) flower. Cultivated in glasshouse, Surrey, England, UK
Scarlet monkey flower (Mimulus cardinalis). The high position of the stamens and stigma ensure that the heads of hummingbird pollinators make contact with them whilst nectaring
Ivy leaved toadflax (Cymbalaria muralis) flowers with orange nectar guides. Surrey, England, UK. April
Bumblebee scarab beetles (Pygopleurus humeralis) feeding on Phelypaea flower (Phelypaea coccinea). Flower has marks that mimic beetle elytra wings to attract pollinators
White dead nettle (Lamium album) with stamens protected from rain by overhanging upper lip. Pollinated by bumblebees. Naturalised in garden, Surrey, England, UK. May
Balsam (Impatiens bicaudata) flowers with lines leading to nectaries. Cultivated. Native to Madagascar
Red trailing bauhinia (Bauhinia kockiana), flowers change from yellow to red with age. Pollinated by butterflies. Cultivated in garden. Native to Malaysia
Ant (Formicidae) nectaring on Geraldton waxflower (Chamelaucium uncinatum). Western Australia Botanic Garden, Kings Park, Perth, Australia. November
Fairy fern (Azolla filiculoides) and Water forget me not (Myosotis scorpioides) in frozen garden pond. Surrey, England, UK. January
Southern double-collared sunbird (Cinnyris chalybeus), male nectaring on Pincushion protea (Leucospermum cordifolium). Cape, South Africa. August
Red wattlebird (Anthochaera carunculata) nectaring on Grevillea (Grevillea preissii). Western Australia Botanic Garden, Kings Park, Perth, Australia. November
Wildflower drifts on disused railway line, flowers include Rosebay willowherb (Epilobium angustifolium), Yellow loosestrife (Lysimachia vulgaris) and Oxeye daisy (Leucanthemum vulgare)
Mandrinette (Hibiscus fragilis), cultivated in breeding program at Kew Gardens, London, UK. Endemic to Mauritius
Gold-tailed melitta bee (Melitta haemorrhoidalis), two males sleeping at night in Peach-leaved bellflower (Campanula persicifolia). In garden, Surrey, England, UK. Focus stacked
Chilean bellflower (Crinodendron hookerianum) releasing pollen via sonication which replicates buzz pollination. Native to Chile. Controlled conditions
Flannel bush (Fremontodendron California Glory ) exposed and overflowing nectaries fluorescuing in UV light. Native to southwest USA
New Zealand tree fuchsia (Fuchsia excorticata) with blue pollen and yellow tipped stigma. Focus stacked
Bloody bellflower (Nesocodon mauritianus), female phase with red nectar surrounding stigma. Native to Mauritius. Cultivated at Kew Gardens, London, England, UK
Goat willow (Salix caprea) female catkin in UV light, fine hairs and nectar fluroescing. Surrey, England, UK. Focus stacked
Southern magnolia / Bull bay (Magnolia grandiflora) flower. The opening and closing of petals controls the time beetle pollinators enter and leave the flower
Malachite sunbird (Nectarinia famosa) male nectaring on King protea (Protea cynaroides). Cape Province, South Africa. August
Texas bluebonnets (Lupinus texensis) and Indian paintbrush (Castilleja coccinea) amongst wildflowers in ditch beside byway. Rains triggered germinaton of annual seeds. Cuero, Texas, USA. April
Winter aconite (Eranthis hyemalis) with ring of nectaries below stamens. Focus stacked
Tulip tree (Liriodendron tulipifera) flower, pollen and nectar fluorescing in UV light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 2/2
Foxglove (Digitalis purpurea), blue nectar guides and anthers, pollen and stigma fluorescing in UV light. Focus stacked
Southern catalpa (Catalpa bignonioides) flowers. Recently opened with yellow nectar guides, purple in older pollinated flowers. Surrey, England, UK. Native to USA
Great yellow bumblebee (Bombus distinguendus) nectaring on Kidney vetch (Anthyllis vulneraria). RSPB Balranald Nature Reserve, North Uist; Outer Hebrides, Scotland, UK. July
Broomrape (Pedicularis longiflora) in wet meadow amongst Horsetail (Equisetum sp). Maroon marks on flowers act as Nectar guides. Road from Barkham to Ruo Er Gai, Sichuan Province, China. July
Sesame (Sesamum indicum) flowers and green unripe pods. Sichuan Province, China
Red poppywort (Meconopsis punicea), raindrops on flower and stem. Medicinal plant in Tibet. Balang Shan, Sichuan Province, China. June
Primula (Primula sikkimensis), Himalayan cowslip (Primula secundiflora) and Crucifer (Brassicaceae) at edge of stream. Kanding, Sichuan Province, China
Sacred lotus (Nelumbo nucifera) seeds embedded in pits in fleshy receptacle. Seeds and rice are made into porridge. Puzhehei National Wetland Park, Yunnan Province, China
Alexanders rhubarb (Rheum alexandrae) and Dwarf purple rhododendron (Rhododendron impeditum) in wet valley. Large bracts on rhubarb protect tiny flowers from UV rays
Paraquilegia (Paraquilegia microphylla) growing in rock fissures with crustose lichen. On cliff, Baimashan, Sichuan Province, China
Sesame (Sesamum indicum), dried seed pods ready for seed harvesting. Sichuan Province, China
Chinese meadow-rue (Thalictrum delavayi) flowers. Male phase below with stigmas hanging below immature stigmas, female phase above after stamens have fallen
Climbing rose (Rosa filipes) flowering in misty forest. Wolong, Sichuan Province, China
Forest on shore of Temnik river, in spring with Siberian Rhododendron (Rhododendron dauricum) and Birch (Betula sp.) Temnik River, Baikal Nature Reserve, Buryatia, Siberia, Russia, May 2015
Springtail (Dicyrtomina ornata) on oak tree leaf, Niedersechsische Elbtalaue Biosphere Reserve, Elbe Valley, Lower Saxony, Germany, December
Mauritian bloody bellflower (Nesocodon mauritianus), female phase with red nectar at base of style. Cultivated at Kew Gardens, London, England, UK. Native to Mauritius
Trailing fuschsia (Fuchsia procumbens) cultivated in alpine house, Surrey, England, UK. Native to New Zealand
Alexanders rhubarb (Rheum alexandrae) and Dwarf purple rhododendron (Rhododendron impeditum) in wet valley with low cloud. Large bracts on rhubarb protect tiny flowers from UV light
Clashach cove surrounded by flowering Gorse (Ulex europaeus) Hopeman, Scotland, UK, May
Great horned owl (Bubo virginianus) preening two chicks in nest in Saguaro (Carnegiea gigantea) cactus, Sonoran desert, Arizona, USA
Beech trees (Fagus sylvatica) at night in winter, with lamp post illuminating the woodland. Felbrigg Woods, Norfolk, England, UK. January
Impression of Birch tree (Betula pendula) in autumn, Scotland, UK. November
