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Small tortoiseshell (Aglais urticae) butterfly flying amongst Scabious (Knautia arvensis) in hay meadow. Wensum Valley, Norfolk, England, UK. July
Cowslip (Primula veris) and habitat, Klein Schietveld, Brasschaat, Belgium. March
Wasp walking around a Spanish love-in-a-mist (Nigella hispanica) flower to sip nectar from ring of blue nectaries, thorax
Hibiscus (Hibiscus sp) flower with stigmas and pollen covered stamens. Focus stacked
Orange bush monkeyflower (Mimulus aurantiacus). Native to California and Oregon, USA. Focus stacked
Peach leaved bellflower (Campanula persicifolia) dissection, petals removed to show how stamens transfer pollen to style via secondary pollen presentation in bud stage. Focus stacked
Rhododendron (Rhododendron sp) flower, anthers and red stigma visible. Garden in Surrey, England, UK
Alpine rose (Rhododendron ferrugineum) in montane grassland above treeline. Giau Pass / Passo di giau, Dolomites, Italy. July 2011
Balsam (Impatiens rosulata) with yellow nectar guides. Endemic to Kitulo Plateau, Tanzania
Allegheny spurge (Pachysandra procumbens) leaves emerging amongst flowers in garden. Flowers emit spicy fragrance. Surrey, England, UK. Native to Asia
Spiderplant (Cleome hirta) flowerhead. Patterns on two banner petals serve as nectar guidesSpiderplant (Cleome hirta) flowerhead.Patterns on two banner petals serve as nectar guides. Tanzania
Rock jasmine (Androsace sericea) flowers. Colour of corona ring indicates age and nectar content of flower. Mountains of Heaven / Tian Shan, Kazakhstan. June
Evening primrose (Oenothera glazioviana) flowering at dusk in garden, Surrey, England, UK. August
Winter aconite (Eranthis hyemalis) flower and bud. Ring of nectaries formed from petals. In garden, Surrey, England, UK. February
Thorowax (Bupleurum rotundifolium) umbel with green bracts, backlit in studio
Masterwort (Astrantia sp) with pink bracts and tiny florets. Cultivated in garden, Surrey, England, UK. Controlled conditions
Foxglove (Digitalis purpurea) flower, backlit with stamens and style visible through corolla. Dark spots on lower lip attract bumblebee pollinators. Pollinated by bumblebees with long proboscis
Wreath nasturtium (Tropaeolum polyphyllum) flowering amongst rocks. Paso Vergara, Chile. December
Bumblebee (Bombus sp) nectaring on Himalayan balsam (Impatiens glandulifera). Surrey, England, UK. September
Love-in-a-mist (Nigella damascena) stamens fluroescing in UV light. Ring of nectaries are dark red with blue UV reflective hairs
Bumblebee (Bombus sp) and Honey bee (Apis mellifera) nectaring on Love-in-a-mist (Nigella damascena) flower. Anthers transferring pollen onto bee backs
Orange-petaled passionflower (Passiflora aurantia) changing colour from cream to orange. Cultivated in glasshouse, Kew Gardens, London, England, UK. Native to New Guinea and northeast Australia
Monarch butterflies (Danaus plexippus), three nectaring on Poinsettia (Euphorbia pulcherrima), resting on red bracts. In butterfly house, Xishuangbanna, Yunnan Province, China
Red bowled anemone (Anemone bucharica) with dark stamens and patterns on petal bases to attract Scarab beetle (Eulasia sp) pollinators
Christmas rose (Helleborus niger) flower, pollen on dehisced anthers and on petals fluorescing in UV light. Nectar ring appearing red. Focus stacked
Blue passionflower (Passiflora caerulea). Flower is pollinated by larger bees. Focus stacked
Large-flowered evening primrose (Oenothera glazioviana). Bulls eye in centre of flower fluorescing in UV light guides pollinators to nectaries. At night in garden. Surrey, England, UK
Spider flower (Cleome sp) with long gonophores. View from above. Cultivated in garden, Surrey, England, UK. Native to South America
Silver Y moth (Autographa gamma) nectaring on Evening primrose (Oenothera glazioviana) with proboscis extended. At night in garden. Surrey, England, UK. August
Honey bee (Apis mellifera) with pollen grains on back. Inside male Squash (Cucurbita sp) flower. Cultivated in garden, Surrey, England, UK. September
Common bee fly (Bombylius major) nectaring on Primrose (Primula vulgaris). This insect is the main pollinator of Primrose. In nature reserve, Surrey, England, UK, April
Hazel (Corylus avellana) catkins dispersing pollen with female flower above on branch. Controlled condition.s Surrey, England, UK
Tulip tree (Liriodendron tulipifera) flower with orange band to attract bees, produces copious nectar. In visible light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 1/2
Foxglove (Digitalis purpurea) corollas with spatter mark nectar guides. Pollinated by long tongued bumblebees able to reach nectar
Seville orange (Citrus u aurantium) flower. Cultivated in glasshouse, Surrey, England, UKSeville orange (Citrus × aurantium) flower. Cultivated in glasshouse, Surrey, England, UK. Focus stacked. Sequence 1/2
Seville orange (Citrus u aurantium) flower fluorescing in UV light, colour attractiveSeville orange (Citrus × aurantium) flower fluorescing in UV light, colour attractive to pollinators. Cultivated in glasshouse, Surrey, England, UK. Focus stacked. Sequence 2/2
Sacred lotus (Nelumbo nucifera) in visible light. Cultivated in glasshouse, Surrey, England, UK. Sequence 1/2
Wasp nectaring on Marsh marigold (Caltha palustris), picking up pollen on head, thorax and hairs on body. Turkey. May
Narcissus fly (Merodon equestris) male nectaring on Rock rose (Cistus x hybridus) in garden, Surrey, England, UK
Bee beetle (Trichius fasciatus), a bee mimic, nectaring on Umbel-flowered catchfly (Silene compacta). Picks up pollen on hairy body. Russian Caucasus. June
Carpenter bee (Xylocopa caffra) male robbing nectar from Ninepin heath (Erica mammosa), not pollinating flower. The Cape, South Africa. February
Rhododendron (Rhododendron souliei) and Leopard plant (Ligularia sp) on bank of river flowing through forest. South of Kangding, Sichuan Province, China. June
China glory flower (Clerodendrum bungei) fruits above red calyxes. Eaten and spread by birds. Tangjiahe National Nature Reserve, Sichuan Province, China
Bird flower (Polygala arillata) for sale as food, Tengchong market, Yunnan Province, China
Handkerchief / Dove tree (Davidia involucrata) inflorescence, stamens with green ovary and multi--rayed style beneath. Native to Sichuan, northwest Yunnan, Guizhou and Hubei, China
Great eggfly (Hypolimnas bolina) butterfly male nectaring on Rough-leaved hydrangea (Hydrangea aspera). Male is highly territorial chasing other insects away
Raindrops on Sacred lotus (Nelumbo nucifera) lily pad. Micro nanostructures create an ultrahydrophobic surface so water slides off leaf. Puzhehai National Wetland Park, Yunnan Province, China
Incarvillea (Incarvillea compacta) with raindrops on petals. Near Zhedou Shan Pass, Sichuan Province, China
