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Dalmatian, head portrait, male aged 4 years
Arctic tern (Sterna paradisaea) in flight, calling, Farne Islands, Northumberland, UK, June
Silhouette of Black-winged stilt (Himantopus himantopus) standing on exposed rock over water, with reflections, Lesbos, Greece
Portrait of an Arctic snow hare (Lepus arcticus groenlandicus) North East Greenland. March
English Oak (Quercus robur) silhouetted on horizon at sunrise. West Milton, Dorset, England, UK, February
Great crested grebes (Podiceps cristatus) performing their elaborate courtship ritual at dawn, Cheshire, UK November
Mute swan (Cygnus olor) in late evening light, Fife, Scotland, UK, November. Photographer quote: The last rays of winter sunshine caught this mute swan as it emerged from a reedbed