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Line of Leaf-cutter ants (Atta sp) carrying leaves, digital composite
Twin Birch (Betula) trees highlighted snow. Senja, Norway, February
Tree sparrow (Passer montanus) feeding a fledgling on a branch of Hawthorn (Crataegus monogyna) blossom, Perthshire, Scotland. June 2013
Barbary macaque (Macaca sylvanus), two adults with baby, overlooking town and ships in sea at sunset. Gibraltar Nature Reserve, Gibraltar. August 2018
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea in the late afternoon, close to Little Brother Island
Humpback whales (Megaptera novangliaea) pod engaged in social foraging by herding herring and other fish with bubble net feeding technique, Chatham Strait, Alaska, USA July
Ants (Formicidae) attacking newly emerged cicada (Cicadacae), Yasuni National Park, Ecuador. Highly commended in the Invertebrates Category of the Wildlife Photographer of the Year Awards (WPOY)
Weaver ants (Oecophylla smaragdina) holding leaves together during nest building, Malaysian Borneo
Bull sharks (Carcharhinus leucas) in Beqa (Benga) Lagoon, a reserve where sharks are fed as part of a commercial diving operation. Viti Levu, Fiji, South Pacific
Coalition of male Lions (Panthera leo) in grassland before storm, Masai-Mara game reserve, Kenya
Blue shark (Prionace glauca) close up, Azores, Portugal. Finalist in Wildlife Photographer of the Year Awards (WPOY) Competition 2014
Harlequin sweetlips (Plectorhinchus chaetodonoides) with cleaning Bluestreak cleaner wrasse (Labroides dimidiatus) and Bicolour cleaner wrasse (Labroides bicolor) South Atoll, Tubbataha Atolls
Galapagos Marine Iguana (Amblyrhynchus cristatus) with Lava Lizard (Trpoidurus) on head. Fernandina island, Galapagos
Three Atlantic sailfish (Istiophorus albicans) hunting cooperatively to force a school of Spanish sardine (Sardinella aurita) into a tight baitball and trapping them at the surface, Isla Mujeres
Red knots (Calidris canutus) in winter plumage feeding co-operatively on tidal mudflats. Holy Island (Lindisfarne), Northumberland, England, UK December
Aerial view of Galapagos sea lion (Zalophus wollebaeki) hunting cooperatively by driving Amberstripe scad fish (Decapterus moruadsi) from open sea to small cove
Galapagos sea lion (Zalophus wollebaeki) hunting cooperatively by driving Amberstripe scad fish (Decapterus moruadsi) from open sea to small cove
Honey bees (Apis mellifera) forming living bridge, Kiel, Germany, June
Leafcutter ants (Atta sp) colony harvesting pieces of a banana leaf, Costa Rica
Silvertip sharks (Carcharhinus albimarginatus) in open water, Revillagigedo islands, Mexico. Pacific Ocean
Greenfinch (Carduelis chloris) pair, one perched on branch and one hovering in snowfall, Derbyshire, February
RF- Bluebells (Hyacinthoides non-scripta / Endymion scriptum) flowering in Broad buckler fern (Dryopteris dilatata), Hallerbos, Belgium. April
Bohemian waxwings (Bombycilla garrulus) feeding on berries, Uto Finland October
Honeybee workers exchanging food - known as trophallaxis (Apis mellifera) Sussex, UK
Bigscale soldierfish (Myripristis berndti), Sweetlips, Philippine chromis with Blueline snappers in background. Komodo National Park, Indonesia
