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Gannet (Morus bassanus) colony on cliffs, viewed from above, Shetland Islands, Scotland, UK, July
Gannet nesting colony on Bass Rock, UK
Little gull (Larus / Hydrocoloeus minutus) Cromer, Norfolk, UK, August
Eider (Somateria mollissima) male in flight in evening light. Aberdeenshire, Scotland, UK, March
Northern gannet {Morus bassanus} head portrait, Bass rock, Scotland, UK
Gannet (Morus bassanus) on coastline, with flowering Sea thrift (Ameria maritima) in foreground, Yorkshire, England, UK. May
Northern gannets {Morus bassanus} on promontary, Noss Island, Bressay, Shetland Islands, Scotland, UK
Looking down cliffs onto Northern gannet (Morus bassanus) colony, Noup of Noss Nature Reserve, Shetland Islands, Scotland, UK
Sandymouth Bay at low tide at sunset, north Cornwall, UK. July 2009
Kimmeridge Bay, Isle of Purbeck, Jurassic Coast World Heritage Site, Dorset, England, UK. Sunset, November 2008
Detail of geological rock formations in the sandstone cliffs of Jaizkibel Beach, Gipuzkoa, Basque Country, Spain, May 2012
Coconut / Robber crab (Birgus latros) climbing coconut tree, Aldabra Seychelles
Green Aurora borealis over the Beaufort Sea, seen from the 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska, October 2011
Killer Whale / Orca (Orcinus orca) large bull swimming in Resurrection Bay, Kenai Fjords National Park, outside Seward, Alaska, May
Long-tailed / Crab-eating macaque (Macaca fascicularis) foraging on coastline at low tide, Bako National Park, Sarawak, Borneo, Malaysia
Grey seal (Halichoerus grypus) on beach with wind blowing sand, Donna Nook, Lincolnshire, UK, November 2008
King Penguin (Aptenodytes patagonicus) male and female rear view looking out over colony, Saint Andrews Bay, South Georgia
Perito Moreno Glacier, panoramic view, Argentina, January 2010
Killer WHale / Orca (Orcinus orca) bull and cow in Resurrection Bay, Kenai Fjords National Park, outside Seward, Alaska, May
Whiskered auklet (Aethia pygmaea), Iony Island / Jonas Island, Sea of Okhotsk, Far East Russia, July
Malachite kingfisher (Alcedo cristata) perched on rush, Intaka Island, Cape Town, South Africa, February
Cardiff Bay in the evening, with meadow flowers on the Barrage. Cardiff, Wales, July 2011
Sunset over Loch Linnhe, Port Appin, nr Oban, Argyll, Scotland, UK
Whitsunday Islands, aerial view, Great Barrier Coral Reef, Queensland, Australia, October 2011
Twelve Apostles rock formations, Great Ocean Road, Port Campbell National Park, Victoria State, Australia, September, 2011
Corcovado Hill and the Christ Redemptor statue in the Atlantic Rainforest of Tijuca National Park, Rio de Janeiro City, Rio de Janeiro State, Southeastern Brazil, August 2011
Mourtia beach and bay with the southeast tip of Samos and Mount Mycale in Turkey sMourtia beach and bay with the southeast tip of Samos and Mount Mycale in Turkeys Dilek Peninsula National Park in the background, east coast of Samos, August 2011
A pod of Bottlenose dolphins (Tursiops truncatus) swimming into the Sado river, Portugal, July
Iceberg off the Antarctic Peninsula, Antarctica, February 2009, Taken on location for BBC Frozen Planet series
View from Bieton Hill looking southwards over Mission Beach, near Innisfail, Queensland, Australia, October 2010
Lighthouse on Fanad Head at sunset, County Donegal, Republic of Ireland, UK, August 2011
Beach and cliffs east of Ballintoy village, North Antrim, Northern Ireland, UK, July 2011
Lapwing (Vanellus vanellus) group roosting in the lagoons at Titchwell RSPB Reserve, Norfolk, UK, October. Non-ex
Ponta de Sao Lourenco Nature Reserve, Madeira
The isthmus between Greater and Little Sark, bound in snow. The heaviest snowfall on Sark in living memory. Channel Islands, UK, December 2010
A view of a forested inlet lit by low sunlight. Aggro Beach, west coast of Vancouver Island, Canada, September 2010
Driftwood on a beach with the coastline stretching to the horizon. Wickaninnish Beach, Pacific Rim National Park Reserve, West coast of Vancouver Island, Canada, September 2010
Dramatic seas crashing onto a rocky shore. The Broken Group Islands from The Wild Pacific Trail, Ucluelet, West coast of Vancouver Island, Canada, September 2010
Hikers crossing a thin footbridge over a river. The West Coast Trail, Pacific Rim National Park, Vancouver Island, Canada, September 2010, model released
Dramatic light on Benijo Beach. Anaga Peninsula, Tenerife, Canary Islands, February 2011
Cliffs and sea-stacks on the coast of El Hierro Island. Canary Islands, February 2011Cliffs and sea-stacks on the coast of " El Hierro" Island. Canary Islands, February 2011
Panther Chameleon {Furcifer pardalis} climbing along branch on beach, Masoala Peninsula National Park, north east Madagascar
Gannet colony at Muriwai, North Island, New Zealand
Group of South American fur seals (Arctocephalus australis) swimming near the rocky shore. Puerto Deseado, Patagonia, Argentina, March 2008
Killer whale / Orca (Orcinus orca) surfacing with three seabirds flying, Tysfjord, Norway, November
Aerial view of Dyrholaey and Reynisdrangar, near Vik, in South Iceland, April 2010
Laysan albatross (Phoebastria immutabilis) head portrait calling mate, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, March
Northern elephant seal (Mirounga angustirostris) close-up of weaner / juvenile sleeping, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, April