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Seaweeds including Thongweed / Sea thong (Himanthalia elongata), Tangleweed kelp (Laminaria digitata), Toothed wrack (Fucus serratus) and Dulse (Palmaria palmata)
Rock pipit (Anthus petrosus) foraging for invertebrates among seaweed on the strand line as the tide rises at sunset, Cornwall, UK, September
Otter (Lutra lutra) portrait on shore with boats in background. Argyll, Scotland, UK, August
Otter (Lutra lutra) feeding, Argyll, Scotland, UK, August
Yellow horned poppy (Glaucium flavum) growing on shingle beach, Norfolk, UK, August
Dog walkers walking along The Wash in evening low tide, Norfolk, UK August 2013
Chocolate working cocker spaniel amongst sand dunes. Formby, Merseyside, UK
Sea stacks at Mangurstadh / Mangersta Beach, Isle of Lewis and Harris, The Outer HebridesSea stacks at Mangurstadh/Mangersta Beach, Isle of Lewis and Harris, The Outer Hebrides, Scotland, UK. October
Luskentye beach, mountains and incoming tide, Isle of Lewis and Harris, Outer Hebrides, Scotland, UK. October 2018
Bamburgh Castle and sand dunes, late evening light, Bamburgh, Northumberland, UK. March 2018
Godrevy Lighthouse at dusk, St. Ives Bay, West Cornwall, England, UK. February 2016
RF - Hartland Quay, late evening light, Hartland, Devon, UK. July
Northern gannets (Morus bassanus) performing sky-pointing display with outspread wings as part of courtship, Cape St. Marys Ecological Reserve, Newfoundland, Canada. June
Puffin (Fratercula artica) with orange sea lichen (Caloplaca marina) covered roch behind, Shiant Isles, Outer Hebrides, Scotland, UK. June
Feral sheep with Galtachan islands behind, Shiant Isles, Outer Hebrides, Scotland, UK. June 2018
Razorbill (Alca torda) orange sea lichen (Caloplaca marina) covered roch behind, Shiant Isles, Outer Hebrides, Scotland, UK. June
Mangroves on Tupeti Island, Southern Lagoon, Forgotten Coast, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Coral formations at low tide in Prony Bay in the Southern Lagoon, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Rock flagtail (Kuhlia rupestris) fish in river on the Southern Lagoon, Forgotten Coast, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Griffon vulture (Gyps fulvus) landing on coast next to flock. Cantabria, Spain, March
Greater flamingos (Phoenicopterus roseus) flock, Sado Estuary, Portugal. January
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Lemon shark pups (Negaprion brevirostris) spend the first 5-8 years of their life in mangrove forests. The tangle of roots provides protection from predators
Part of a submerged Holocene forest, revealed after a winter storm from beneath the sand at Hells Mouth (Porth Neigwl), Abersoch Wales, UK March 2014
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Killer whale (Orcinus orca) hunting for herring fish (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Killer whale (Orcinus orca) hunting inside herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Common eider (Somateria mollissima), female on her nest with sea behind, Finland, May
Southern Rockhopper penguin (Eudyptes chrysocome) colony, Kidney Island, Falkland Islands, October. Vulnerable species
Silver Drummer (Kyphosus sydneyanus) fish off the coast of Neds Beach, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Southern elephant seal (Mirounga leonina), fight between two males, Saint Andrew, South Georgia
Dingo (Canis lupus dingo) on the beach at night, Fraser Island UNESCO World Heritage Site. Queensland, Australia, November
Winter storm on the coast near Arnastapi (rock arch of Gatklettur), Snaefellsness Peninsula, Iceland, February 2016
Aerial view of Cape Town city with Table Mountain, South Africa, taken from helicopter, May 2011
Shag (Phalacrocorax aristotelis) two heads showing above rock, Vardo, Norway March
Killer whale (Orcinus orca) adult male surfacing at dusk surrounded by birds, who watch for herrings (Clupea harengus) left after they have fed, Andenes, Andoya island, North Atlantic Ocean, Norway
Great white egrets (Casmerodius albus) reflected in Pulicat Lake, Tamil Nadu, India, January 2013
Marine iguana (Amblyrhynchus cristatus) in the surf, Black Beach, Floreana Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) in water, Puerto Egas, Santiago Island, Galapagos
Galapagos lava lizard (Microlophus albemarlensis) sitting on Marine iguana (Amblyrhynchus cristatus) Punta Espinosa, Fernandina Island, Galapgos
Princely fiddler crab (Uca princeps), Bahia Magdalena, Baja California Peninsula, Mexico, June
Eggs of Common whelk (Buccinum undatum) on seaweed washed up on beach, Sark, British Channel Islands
Pennants Topshell (Gibbula pennanti) and Purple Topshell (Gibbula umbilicalis) sea shells on beach, showing difference between species, sea shells on beach, Sark, British Channel Islands
Hairy Crab (Pilumnus hirtellus) on seaweed on the beach, Sark, British Channel Islands
Eggs of Long-spined sea scorpion (Taurulus bubalis) on sea shore, possibly being predated by Painted Topshell, Guernsey, British Channel Islands
Gatklettur, Arnarstapi, Snaefellsnes Peninsula, Iceland, March 2017
Bamburgh Castle, Northumberland, England, UK, March 2009
Sand patterns with ice between ripples of the sand, front of Burnham-on-Sea Lighthouse in winter, Somerset, England, UK, January.. January 2010