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Santa Fe land iguana (Conolophus pallidus) next to Tree Prickly pear (Opuntia echios) Santa Fe Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) on rock, with pounding waves, Puerto Ayora / Academy Bay, Santa Cruz Island, Galapagos
Great blue heron (Ardea herodias) with hatchling sea turtle prey, Las Bachas, Santa Cruz Island, Galapagos
Galapagos sea lion (Zalophus wollebaeki) hunting, Champion Islet, Floreana Island, Galapagos
Nazca booby (Sula granti) on coast, with Vampire ground finches (Geospiza septentrionalis). These finches peck at birds, feeding on blood when other food sources are scarce, Wolf sland, Galapagos
Galapagos fur sea (Arctocephalus galapagoensis) pup, curiously approaching group of Marine iguanas (Amblyrhynchus cristatus) Cape Hammond, Fernandina Island, Galapagos
Wildlife on the coast of Cape Hammond including Galapagos flightless cormorant (Phalacrocorax harrisi), Sally lightfoot crab (Grapsus grapsus) and Marine iguana (Amblyrhynchus cristatus)
Nazca booby (Sula granti) in flight at dusk. Punta Suarez, Espanola Island, Galapagos
Green turtle (Chelonia mydas) female digging nest, North coast, Santa Cruz Island, Galapagos
Saline lagoon bordered by salt bush, with typical arid vegetation beyond, Rabida Island, Galapagos Islands, June 2011
California least terns (Sternula antillarum browni) courtship behaviour, Bolsa Chica Ecological Reserve, California, USA June
Thatch palm (Thrinax radiata), Guanahacabibes Peninsula National Park, Pinar del Rio Province, western Cuba
Cuban / Clouded rock iguanas (Cyclura nubila), on beach, Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba
Desmarests hutia (Capromys pilorides doceleguas) on beach, Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba
Orbiculate cardinalfish (Sphaeramia orbicularis) sheltering amongst roots of mangroves (Rhizophora sp.) in Raja Ampat, West Papua, Indonesia. Pacific ocean
View of coastal beach with palm trees in Raja Ampat, West Papua, Indonesia. Pacific Ocean
Grey seal (Halichoerus grypus), mother and young interacting on beach, UK
Galapagos Sea Lion (Zalophus californianus wollebacki) resting on rocks at sunset on North Seymour Island, Galapagos, Ecuador. December
Arctic fox (Alopex lagopus). Hornstrandir, Iceland. Blue colour morph in winter coat. Photographed by camera trap. March
RF - Bar-tailed godwit (Limosa lapponica) in flight over the Wash, Norfolk, UK. November. (This image may be licensed either as rights managed or royalty free.)
Pied Oystercatcher (Haematopus longirostris) pair, Crab Island, Torres Strait Islands, Queensland, Australia
Bottlenose dolphin (Tursiops truncatus) near surface, Sado Estuary, Arribida, Portugal
Coastal landscape with empty beach Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, July
Little Tern (Sterna albifrons) chick walking back to parent sitting on nest scrape on beach. Gronant Dunes, Denbighshire, Wales, UK, June
Atlantic puffin (Fratercula arctica) in breeding plumage flapping wings on sea cliff top in seabird colony, Scotland, UK, May. Digital composite
Great skua (Stercorarius skua) in flight and spectacular coastline with sea stacks and cliffs, home to breeding sea birds at Hermaness, Unst, Shetland Islands, Scotland. May 2018. Digital composite
St Michael's Mount at sunrise with a rainbow over Penzance, viewed from Marazion, West Cornwall, UK. February 2019
Otter (Lutra lutra) family at rest on seaweed covered rock, taken in the Inner Hebrides, Scotland, UK, November. Highly commended in BWPA Competition 2019
Coast of l Ile-des-Pins with the New Caledonia pines (Araucaria columnaris) that gave the name to the island at sunset, New Caledonia, Pacific Ocean
Mediterranean gull (Larus melanocephalus) in summer plumage, Hayling Island, Hampshire, England, UK, July
African jackass penguins (Spheniscus demersus) three heading to sea in the early morning to forage for food, Cape Town, South Africa, Endangered IUCN
Atlantic walruses (Odobenus rosmarus) resting on ice, with two large individuals facing off just before the one on the right stabbed the other with his tusks, Svalbard, Norway, June
Dwarf willows add color along a small glacial stream before icebergs in Hare Fjord, Scoresby Sund, Greenland, August
Brydes whale (Balaenoptera brydei) fin at surface in coastal waters, volcano in background. Punta Rasca, Tenerife, Canary Islands
Desmarests hutia (Capromys pilorides) in tree on beach. Gardens of the Queen National Park, Cuba
Qinghai Lake shore with Great cormorants (Phalacrocorax carbo sinensis) colony nesting on rocks. Qinghai Province, China. May 2009
Bedding planes and joint network in Jurassic age Blue Lias limestone and shale exposed at low tide at Nash Point, South Glamorgan Heritage Coast, Wales, September 2017
Several faults cutting through Jurassic age, Liassic Limestones (Blue Lias) near Southerndown, South Glamorgan Heritage Coast, Wales, UK. September
Autumnal landscape on Porlock Hill with Dunkerry Beacon beyond, Exmoor National Park, Somerset, England, UK, November 2014
Starlings (Sturnus vulgaris) resting on Blackpool Beach, UK. October
Espanola mockingbird (Mimus macdonaldi) on beach, Galapagos
Little tern (Sterna albifrons ) feeding sand eel (Hyperoplus spp) to young chick. Gronant Dunes, Denbighshire, Wales, UK, June
Sally lightfoot crabs (Grapsus grapsus) on the beach at Puerto Egas, Santiago Island, Galapagos, May
Flock of wading birds flying over The Wash, at dusk, Norfolk, UK, September 2008
Corfe castle and village at dawn with mist, Corfe Castle, The Purbecks, Dorset, UK. September 2014
Grey seal (Halichoerus grypus) pup on a breeding beach, Orkney, Scotland, UK, April
Grey seal (Halichoerus grypus) huge group hauled out on a beach, Island of Mingulay, west coast of Scotland. May
Route One, Southeast Iceland. December