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Western Diamondback Rattlesnake (Crotalus atrox), adult in striking pose in wildflower field. Laredo, Webb County, South Texas, USA, April
Black Swallowtail Butterfly (Papilio polyxenes), female on Cowpen Daisy / Golden Crownbeard (Verbesina encelioides). Laredo, Webb County, Texas, USA, April
Red Fox (Vulpes vulpes) pouncing in grass. Vosges, France, July
Roe Deer (Capreolus capreolus) swimming across River Allier, France, February
Blue Tit (Parus caeruleus) perched on mossy branch. Breton Marsh, French Atlantic Coast, November
Shipwreck of the small freighter Ora Verde that sank in 1980, Grand Cayman Island, Caribbean Sea
Close up of skin of Rainbow boa (Epicrates cenchria cenchria) captive, from French Guiana
White-lipped / Green tree viper (Cryptelytrops albolabris) captive, from SE Asia
Stellers Sealion (Eumetopias jubata) stampede into the water. Clayoquot Sound, Vancouver Island, Canada, March
Portrait of a Common Racoon (Procyon lotor). Stanley Park, Vancouver, Canada, March
Two orphan baby Elephants (Loxodonta africana) being affectionate. David Sheldrick Wildlife Trust Nairobi Elephant Nursery, Kenya, July 2010
Wild horse / mustang, mare rolling, Pryor Mountains, Montana, USA, July
Portrait of a wild Horse (Equus caballus). The foothills of southwest Reno, Nevada, USA, May
Wild stallion (Equus caballus) lying down in the sagebrush and yawning. The foothills south of Reno, Nevada, USA, May
Female Rocky Mountain bighorn sheep ewe (Ovis canadensis canadensis) warning her newborn lamb that a rocky ledge is too close and dangerous
Honey bee (Apis mellifera) approaching a red poppy (Papaver rhoeas) in a small meadow of wildflowers. Nevada, USA, July
Golden eagle (Aquila chrysaetos) about to land on snow, Oulanka NP, Finland, February
Portrait of an Emerald Glass Frog (Espadarana / Centrolenella prosoblepon). Captive. Choco Region of northwest Ecuador
Poets Daffodil (Narcissus poeticus). Montseny Nature Reserve, Barcelona province, Catalonia, Spain, May
Whale Shark (Rhincodon typus), eating krill and plankton at the surface. Gulf of Mexico, Mexico, North America, August
Panther Chameleon {Furcifer pardalis} climbing along branch on beach, Masoala Peninsula National Park, north east Madagascar
Fragrant water lilies (Nymphaea odorata) in Lake Skadar, Montenegro, May 2008
Southern plains grey / Hanuman langur {Semnopithecus dussumieri} two adolescents sitting, Thar desert, Rajasthan, India
Red Deer (Cervus elaphus) male standing in snow. The Netherlands, January
Five Mallard (Anas platyrhynchos) walking in a line across snow. Three males and two females. The Netherlands, January
Grey Heron (Ardea cinerea) leaving footprints while walking through snow. The Netherlands, January
Whooper Swan (Cygnus cygnus) with its beak tucked under its wing. The Netherlands, June
Portrait of Grey Heron (Ardea cinerea) looking directly at lens. The Netherlands, June
Atlantic salmon (Salmo salar) moving upriver to spawn. Lligwy River nr Betws y Coed, Gwynedd, North Wales, UK, November
Reef with Pygmy Sweepers (Parapriacanthus ransonetti), Barrel Sponge (Xestospongia testudinaria), and soft corals. Komodo National Park, Indonesia, October
Close up of feathers of cock Pheasant (Phasianus colchicus)
Apollo butterfly (Parnassius apollo) on thistle head in alpine meadow. Nordtirol, Austrian Alps
Brown Hare (Lepus europaeus) portrait gazing alertly. Perthshire, Scotland, June
Domestic dog, Scottish Terrier / Aberdeen Terrier, studio portrait
Brown bear (Ursus arctos) running through water, fishing, Kamchatka, Far east Russia, August
Brown bear (Ursus arctos) walking beside lake, Kamchatka, Far east Russia, August
Alpine meadows with Caucasian scabious (Scabiosa caucasica) in Bezengi gorge, July, 3000m. W-C Caucasus, Kabardino-Balkarskiy nature reserve, Russia
Brown bear (Ursus arctos) standing beside lake looking for salmon, Kamchatka, Far east Russia, August
Wood Ant (Formica rufa) workers on top of their nest synchronise ejection of formic acid droplets to maximise deterrent effect on potential predator, UK
A young California sealion (Zalophus californianus) playing with a starfish, inside an underwater cave. The pups will pick up a variety of objects (such as starfish, lumps of coral)
South African fur seal (Arctocephalus pusillus pusillus) swimming near surface, Seal Island. False Bay, South Africa, July 2010
Hawaiian goose / Nene (Branta sandvicensis) lookig down, Slimbridge, Gloucestershire, UK, Captive, native to Hawaii, USA, Vulnerable species, September
Killer whale / Orca (Orcinus orca) surfacing with three seabirds flying, Tysfjord, Norway, November
Zanzibar Red Colobus (Procolobus kirkii) head portrait, Jozani Chwaka Bay NP, Zanzibar, Tanzania
Proboscis Monkey (Nasalis larvatus) female sitting in tree, partially obscured by soft focus leaves in foreground, Bako National Park, Sarawak, Borneo, Malaysia
Australian Shepherd, blue-merle, head portrait with autumn leaves behind
Norfolk Terrier portrait sitting in autumn foliage
Solar eclipse 63% on 4 January 2011, seen from Barcelona, Spain