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Atlantic puffin (Fratercula arctica) close up of head, Fair Isle, Shetland, Scotland, UK, July
Atlantic Puffins (Fratercula arctica) bill rubbing, Skomer Island, Wales, UK, April
Atlantic puffin (Fratercula arctica) landing, wings outstretched. Machias Seal Island, off the coast of Maine, USA. July
Puffin (Fratercula arctica) standing on flowering Sea thrift (Armeria maritima). Portrait. Fair Island, Shetland Islands, Scotland, July
Oystercatcher (Haematopus ostralegus) by the coast, Island of Runde, Norway July
Dotterel (Charadrius morinellus) standing on rock, Cairngorms National Park, Scotland, UK, May
Pied avocet (Recurvirostra avosetta) feeding in shallow water at dawn, Etosha National Park, Namibia
Arctic tern (Sterna paradisaea) hovering above the ocean, Svalbard, Norway June
Piping Plover (Charadrius melodus) brooding three chicks with a fourth approaching, northern Massachusetts, USA.June. Endangered species
Puffin (Fratercula arctica) on a cliff edge with flowering sea thrift (Armeria maritima) Great Saltee, Saltee Islands, County Wexford, Ireland, June
Swallow-tailed gull (Creagrus furcatus) Plazas Island, Galapagos
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Black-headed gull (Chroicocephalus ridibundus) in flight, artistically blurred photograph, Cheshire, UK, February. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Atlantic Puffin (Fratercula arctica) one landing among resting group, Sule Skerry, Scotland, UK, July
Atlantic Puffins (Fratercula arctica) pair bill rubbing, part of ritual courtship, Skomer Island, Wales, UK, May
Sanderling (Calidris alba) flock foraging at sunset, Britanny, France, October
Grizzly bear (Ursus arctos horribilis) and Grey wolf (Canis lupus) on beach, Katmai National Park, Alaska, USA, August
The River Thames and Houses of Parliament at Westminster at sunset with gull, UK, November 2005
Caspian Tern (Sterna caspia) in flight, diving for prey. Lake Ellesmere, Canterbury, South Island, New Zealand
Golden plover (Pluvialis apricaria) chick camouflaged in ground nest, Norway June
Woodcock (Scolopax rusticola), in undergrowth in winter, Brittany, France
Pied avocet (Recurvirostra avosetta) adult parent and chick together. Suffolk, UK. June
Puffins (Fratercula arctica) swimming underwater. Puffins spend most of their lives at sea and are excellent underwater swimmers, which is how they catch small fish, their main food
Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May
Small flock of Lapwings (Vanellus vanellus) flying in to rest after migration. Cresswell Pond, Druridge Bay, Northumberland, England, UK, September
Puffins (Fratercula arctica) Shaint Isles, Scotland, UK, June
Green sandpiper (Tringa ochropus) with small worm it has caught in shallow freshwater lake, Gloucestershire, UK, August
Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion
American avocet (Recurvirostra americana), adult in breeding plumage foraging by sweeping its bill from side to side through shallow water, Orange County, California, USA, April
California Gull (Larus californicus), foraging for alkali flies (Ephydra hians) by running through swarm snapping its bill, on the shore at Mono Lake, California, USA. Digital frame expansion
Black-necked stilts (Himantopus mexicanus) pair performing post-copulatory display (as copulation ends, the male slides off the female leaving his wings draped over her)
Black-necked Stilt (Himantopus mexicanus), foraging in water, with reflection, Bear River Migratory Bird Refuge, Utah, USA, May
Black skimmers (Rynchops niger), three in non-breeding plumage on beach, Fort De Soto Park, St. Petersburg, Florida, USA. January. Digitally altered to remove distractions
Black terns (Chlidonias niger), pair on a perch, vocalizing during courtship (termed Erect Posture display), Montezuma National Wildlife Refuge, New York, USA, May
Royal tern (Sterna maxima) perching on a sand castle on a beach, Fort De Soto Park, St. Petersburg, Florida, USA, March
Crested auklets (Aethia cristatella) pair interacting while perched on rock, St Paul Island, Pribilofs, Alaska, USA, July
Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Puffins (Fratercula arctica) and Razorbills (Alca torda) with Eilean Mhuire behind, Shiant Isles, Outer Hebrides, Scotland, UK. June, 2018
Black-headed gull (Chroicocephalus ridibundus) flock flying over a frozen lake in falling snow, Wiltshire, UK, March
Lapwing (Vanellus vanellus) portrait in upland hay meadow, Upper Teesdale, CO Durham, England, UK, June
Common Snipe (Gallinago gallinago) autumn courtship display, Germany, August
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
Razorbill (Alca torda) landing, Great Saltee Island, County Wexford, Ireland, June
Puffin (Fratercula arctica) wings spread backlit, Great Saltee Island, County Wexford, Republic of Ireland, June
Semipalmated sandpiper (Calidris pusilla) flock in flight over water, Johnsons Mills Reserve, Bay of Fundy, New Brunswick, Canada, July
Swallow-tailed gull (Creagrus furcatus) calling, Genovesa Island, Galapagos
Atlantic puffin (Fratercula arctica) portrait, Machias Seal Island, Maine, USA. July
Woodcock (Scolopax rusticola) in snow, portrait, Berwickshire, Scotland, UK. January
Common tern (Sterna hirundo) catching a mayfly, Hjalstaviken, Uppland, Sweden. Nordic Nature Photo Competition 2021 Highly Commended - Nordic Bird Photo of the Year category
Black-headed gull (chroicocephalus ridibundus) diving into water, hunting fish / shellfish at sunrise. Baltic Sea, Poland. Sequence 1 of 2
RF - Atlantic puffin (Fratercula arctica) flying, Hornoya, Varanger, Finnmark, Norway (This image may be licensed either as rights managed or royalty free.)
Eurasian oystercatcher (Haematopus ostralegus) walking over tidal mudflats in low winter sunshine. Holy Island (Lindisfarne) tidal mudflats, Northumberland, England, UK. December
Common tern (Sterna hirundo) flying low over water. Hjalstaviken, Uppland, Sweden
Brown skua (Stercorarius antarcticus), Ronge Island, Antarctica
RF - Atlantic Puffins (Fratercula arctica) flying on to the cliff top and gathering around burrows. Isle of Lunga, Treshnish Isles, Isle of Mull, Scotland
Arctic tern (Sterna paradisaea) adult calling from rock in breeding colony. Isle of May, Scotland, UK, July
Atlantic puffin (Fratercula arctica) taking off, Machias Seal Island, Maine, USA, June
Common tern (Sterna hirundo) close up in flight, Tenerife, Canary Islands
Brown noddy (Anous stolidus) two mobbing Brown pelican (Pelecanus occidentalis) Turtle Cove, Santa Cruz Island, Galapagos
California least terns (Sternula antillarum browni) courtship behaviour, Bolsa Chica Ecological Reserve, California, USA June
Pied Oystercatcher (Haematopus longirostris) pair, Crab Island, Torres Strait Islands, Queensland, Australia
Little Tern (Sterna albifrons) at the nest amongst Black oats (Avena strigosa) North Uist, Scotland, UK, July
Crab-plover (Dromas ardeola) three walking in a row on mudflat, two with food in beaks. Oman, June
Atlantic puffin (Fratercula arctica) swimming on the ocean surface, Svalbard, Norway, June
Brunnichs guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018
Lapwing (Vanellus vanellus) Titchwell RSPB Reserve, Norfolk, England, UK. November
Swallow-tailed gull (Creagrus furcatus) in flight, Plazas Islands, Galapagos
Two Little terns (Sterna albifrons) at the nest amongst Black oats (Avena strigosa) growing on shell rich sands Machair, North Uist, Scotland, UK, June
Little tern (Sterna albifrons) at the nest amongst Black oats (Avena strigosa) growing on shell rich sands Machair, North Uist, Scotland, UK, June
Little tern (Sterna albifrons) with newly hatched chick, Gronant dunes, Denbighshire, Wales, UK, June
Little tern (Sterna albifrons) nest with three eggs in shingle ridge. County Wicklow, Ireland, June
Pectoral sandpiper (Calidris melanotos) juvenile feeding around edge of shallow flood, Llanrhystud, Ceredigion, West Wales, UK, September
Pectoral sandpiper (Calidris melanotos) juvenile, migrant, Llanrhystud, Ceredigion, West Wales
Turnstones (Arenaria interpres) and Purple sandpipers (Calidris maritima) over wintering on the beaches of the Orkney Isles, Scotland, UK, April
Common tern (Sterna hirundo) in flight, Norfolk, UK, August
Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July
Dunlin (Calidris alpina) portrait. Vardo, Finnmark, Norway. May
Turnstone (Arenaria interpres) in summer plumage, on Norfolk shoreline in breaking wave, England, UK, May
American oystercatcher (Haematopus palliatus) adult feeding chick, Long Island, New York, USA. June
American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May
Tufted puffin (Fratercula cirrhata), adult in breeding plumage flying in preparing to land, St. Paul, Pribilof Islands, Alaska, USA, July
Wilsons Phalaropes (Phalaropus tricolor) flock flying over tufa formations at Mono Lake in late summer, with reflection, California, USA. July 2013
Horned Puffin (Fratercula corniculata) flying over a pattern of kelp and sea foam, photographed from above, St. Paul, Pribilof Islands, Alaska, USA. July
Spotted sandpiper (Actitis macularius), with head submerged as it jabs at underwater prey, Caroline, New York, USA, May