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Atlantic puffin (Fratercula arctica) landing, wings outstretched. Machias Seal Island, off the coast of Maine, USA. July
Atlantic puffin (Fratercula arctica) close up of head, Fair Isle, Shetland, Scotland, UK, July
Atlantic Puffins (Fratercula arctica) bill rubbing, Skomer Island, Wales, UK, April
Puffin (Fratercula arctica) standing on flowering Sea thrift (Armeria maritima). Portrait. Fair Island, Shetland Islands, Scotland, July
Swallow-tailed gull (Creagrus furcatus) Plazas Island, Galapagos
Pied avocet (Recurvirostra avosetta) feeding in shallow water at dawn, Etosha National Park, Namibia
Atlantic Puffins (Fratercula arctica) pair bill rubbing, part of ritual courtship, Skomer Island, Wales, UK, May
Arctic tern (Sterna paradisaea) hovering above the ocean, Svalbard, Norway June
Piping Plover (Charadrius melodus) brooding three chicks with a fourth approaching, northern Massachusetts, USA.June. Endangered species
Puffin (Fratercula arctica) on a cliff edge with flowering sea thrift (Armeria maritima) Great Saltee, Saltee Islands, County Wexford, Ireland, June
Oystercatcher (Haematopus ostralegus) by the coast, Island of Runde, Norway July
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Black-headed gull (Chroicocephalus ridibundus) in flight, artistically blurred photograph, Cheshire, UK, February. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
The River Thames and Houses of Parliament at Westminster at sunset with gull, UK, November 2005
Atlantic Puffin (Fratercula arctica) one landing among resting group, Sule Skerry, Scotland, UK, July
Sanderling (Calidris alba) flock foraging at sunset, Britanny, France, October
Golden plover (Pluvialis apricaria) chick camouflaged in ground nest, Norway June
Caspian Tern (Sterna caspia) in flight, diving for prey. Lake Ellesmere, Canterbury, South Island, New Zealand
Puffins (Fratercula arctica) swimming underwater. Puffins spend most of their lives at sea and are excellent underwater swimmers, which is how they catch small fish, their main food
Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May
Small flock of Lapwings (Vanellus vanellus) flying in to rest after migration. Cresswell Pond, Druridge Bay, Northumberland, England, UK, September
Puffins (Fratercula arctica) Shaint Isles, Scotland, UK, June
Green sandpiper (Tringa ochropus) with small worm it has caught in shallow freshwater lake, Gloucestershire, UK, August
Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion
American avocet (Recurvirostra americana), adult in breeding plumage foraging by sweeping its bill from side to side through shallow water, Orange County, California, USA, April
California Gull (Larus californicus), foraging for alkali flies (Ephydra hians) by running through swarm snapping its bill, on the shore at Mono Lake, California, USA. Digital frame expansion
Black-necked stilts (Himantopus mexicanus) pair performing post-copulatory display (as copulation ends, the male slides off the female leaving his wings draped over her)
Black-necked Stilt (Himantopus mexicanus), foraging in water, with reflection, Bear River Migratory Bird Refuge, Utah, USA, May
Black terns (Chlidonias niger), pair on a perch, vocalizing during courtship (termed Erect Posture display), Montezuma National Wildlife Refuge, New York, USA, May
Royal tern (Sterna maxima) perching on a sand castle on a beach, Fort De Soto Park, St. Petersburg, Florida, USA, March
Crested auklets (Aethia cristatella) pair interacting while perched on rock, St Paul Island, Pribilofs, Alaska, USA, July
Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Puffins (Fratercula arctica) and Razorbills (Alca torda) with Eilean Mhuire behind, Shiant Isles, Outer Hebrides, Scotland, UK. June, 2018
Lapwing (Vanellus vanellus) portrait in upland hay meadow, Upper Teesdale, CO Durham, England, UK, June
Common Snipe (Gallinago gallinago) autumn courtship display, Germany, August
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
Razorbill (Alca torda) landing, Great Saltee Island, County Wexford, Ireland, June
Puffin (Fratercula arctica) wings spread backlit, Great Saltee Island, County Wexford, Republic of Ireland, June
Semipalmated sandpiper (Calidris pusilla) flock in flight over water, Johnsons Mills Reserve, Bay of Fundy, New Brunswick, Canada, July
Swallow-tailed gull (Creagrus furcatus) calling, Genovesa Island, Galapagos
Atlantic puffin (Fratercula arctica) in flight, Machias Seal Island off the coast of Maine, USA, July
Ruddy turnstone (Arenaria interpres) in breeding plumage. Yukon Delta National Wildlife Refuge. Alaska, USA. June
Atlantic puffins (Fratercula arctica) amongst flowering Sea campion (Silene uniflora), Skomer Island, Pembrokeshire, Wales, UK. May
Common tern (Sterna hirundo) in flight over water, Hampstead Heath, London, UK, August
Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July
Black-headed gull (Chroicocephalus ridibundus) alighting on water at dawn, Cheshire, UK, November. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Black-bellied Plover (Pluvialis squatarola) chicks in the nest, seen from overhead, Yukon Delta National Wildlife Refuge, Alaska, USA June
Atlantic Puffin (Fratercula arctica) in flight, coming into land with beak full of sand eels (Ammodytes tobianus). Fair Isle, Shetland Islands, Scotland, UK, July