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Gemsbok (Oryx gazella) in sand dunes, Namibia. Finalist in the Mammals Category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
Quiver tree (Aloe dichotoma) with the Milky Way at night, and light pollution from town in the distance, Keetmanshoop, Namibia. Colours accentuated digitally
Pied avocet (Recurvirostra avosetta) feeding in shallow water at dawn, Etosha National Park, Namibia
Chlorophyte or green alga (Volvox aureus) in pond water. UK. The larger spheres are approximately 0.5mm in diameter. Digital composite image
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Crested newt (Triturus carnifex) Gelderland, The Netherlands, April. Winner of the Amphibians category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
Snares island crested penguin (Eudyptes robustus) colony on the coast, high angle view, Snares Island, New Zealand
Wild boar (Sus scrofa) at night with the milky way in the background, Gyulaj, Tolna, Hungary. August. Taken using long exposure with flash at night
Wild boar (Sus scrofa) in old mixed conifer and broadleaf forest, Punia Forest Reserve, Lithuania, May
Leopard (Panthera pardus) cub looking up at birds (out of frame) with mother in background, Khwai River, Botswana
Aerial view of Amazon Rainforest and the Yavari River, and oxbow lake, Peru. July 2015
Golden-mantled tamarin (Saguinus tripartitus) captive
Snow Leopard (Panthera uncia) climbing up mountain slope, Hemis National Park, India, February
Eastern cottontail (Sylvilagus floridanus) Laredo Borderlands, Texas, USA. April
Great crested newt (Triturus cristatus) male in breeding condition, amongst Soft hornwort (Ceratophyllum submersum), Herefordshire, England, UK, May. Captive
Red fox (Vulpes vulpes) peering from behind tree with bluebells in foreground, Cheshire, June
Emperor moth (Saturnia pavonia) female wide angle view showing heather moorland habitat. Peak District National Park, UK. April
Orange Tip Butterfly (Anthocharis cardamines) male resting on Bluebell (Endymion / Hyacinthoides nonscriptus). Peak DIstrict National Park, Derbyshire, UK. April
Leopard (Panthera pardus) hunting Springbok (Antidorcas marsupialis) Etosha, Namibia, Finalist in the Mammals Category of the Wildlife Photographer of the Year 2015
Fork-leaved sundew (Drosera bipinata) leaves unfurling. Cultivated species, occurs in Australia and New Zealand
Wild Garlic / ramsons (Allium ursinum) leaves emerging in early spring. Peak District National Park, Derbyshire, UK. March
A split level image of Southern stingray (Hypanus americanus) swimming over a sand bar. Grand Cayman, Cayman Islands. Caribbean Sea. Digital composite
Caribbean flamingo (Phoenicopterus ruber) about to take off to roosting site, Ria Celestun Biosphere Reserve, Yucatan Peninsula Mexico, January
Portuguese Man-of-War (Physalia physalis) close up of tentacles, Sargasso Sea, Bermuda
Great egret (Ardea alba) flying across lake, Ranthambhore National Park, India
Ribbon sawtail fish (Idiacanthus fasciola) from Atlantic Ocean, at a depth of 800-1000m
Frilled shark (Chlamydoselachus anguineus) specimen with mouth open, from Atlantic Ocean, at a depth of 729m
Large pod of Bottlenose dolphins (Tursiops truncatus) porpoising over waves during annual sardine run, Port St Johns, South Africa
Banded demoiselle (Calopteryx splendens) group of four on plant stems, River Leijgraaf, Nijmegen, the Netherlands, August 2013
Two Great egrets (Ardea alba) standing opposite each other with Grey heron (Ardea cinerea) in between. Lake Csaj, Pusztaszer, Hungary, January
Red-and-green macaw (Ara chloropterus) in flight, Pantanal, Brazil. August
Quiver tree forest (Aloe dichotoma) at night with stars and the Milky Way, Keetmanshoop, Namibia
Eurasian red squirrel (Sciurus vulgaris) reflected in water, Cairngorms National Park, Scotland, United Kingdom, October
View of Le Mont Pourri, with European larch (Larix decidua) forest, Le Monal Vanoise National Park, Savoie, France, October 2015
Big-eye fish (Priacanthus hamrur). Egypt, Red Sea
Bottlenosed dolphin (Tursiops truncatus) porpoising during annual sardine run, Port St Johns, South Africa. June
Lion (Panthera leo) male with scars photographed with side-lit spot light at night. Greater Kruger National Park, South Africa, April
Black rhino (Diceros bicornis) approaching waterhole with her calf close by. Etosha National Park, Namibia
Least chipmunk (Tamias minimus) in snowfall, Grand Teton National Park, Wyoming, USA. October
North American river otter (Lontra canadensis) two in water with autumnal trees reflected in the water, Acadia National Park, Maine, USA, October
Purple gallinule (Porphyrula martinica) in mangrove swamp. Everglades National Park, Florida, USA. March
Snowy egret (Egretta thula) with plumes erect, St. Augustine, Florida, USA, April
Western sandpipers (Calidris mauri) roosting on barrier island tidal flats. Terrebonne Parish, Louisiana, USA. October
Eagle owl (Bubo bubo) in flight through forest, backlit at dawn, Czech Republic, November. Captive
Long-eared owl (Asio otus) perched in tree amongst autumn leaves, Czech Republic, October. Captive
The Crow Stones at sunrise, Peak District National Park, Derbyshire, UK. August 2015
Emerald damselflies (Lestes sponsa) mating, with male above. Peak District National Park, Derbyshire, UK. September
Violet bramble rust (Phragmidium violaceum) causing red colouration on Bramble leaf (Rubus fruticosus aggregate). Peak District National Park, Derbyshire, UK. October