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Nine Stone Close stone circle, Peak District National Park, Derbyshire, UK. November
Horsefly (Philipomyia aprica) Aosta Valley, Monte Rosa Massif, Pennine Alps, Italy. July
Mountain Clouded Yellow (Colias phicomone) profile portrait, Aosta Valley, Monte Rosa Massif, Pennine Alps, Italy
Bohemian waxwings (Bombycilla garrulus) feeding on berries, Uto Finland October
Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June
Golden Eagle (Aquila chrysaetos) portrait in snow, Utajarvi, Finland February
Arctic Tern (Sterna paradisaea) in flight, silhouetted against the moon at dusk, Uto Finland July
Zanzibar red colobus (Piliocolobus kirkii) in tree, Jozany Forest, Zanzibar, Tanzania. Highly endangered
Looking down on the last glaciers near the summit of Mount Kilimanjaro, Tanzania, October 2008
Lobelia (Lobelia deckenii) close up of leaves, alpine species of lobelia that lives on Kilimanjaro, Tanzania
Polar Bear (Ursus maritimus) head portrait with blue tongue sticking out, Svalbard, Norway
White rhinoceros (Ceratotherium simum) mother and calf browsing. Entabeni Safari Conservancy, Limpopo region, Waterberg, South Africa, October
Two Red Foxes (Vulpes vulpes) fighting. The Netherlands. August
Male Marsh pride lion (Panthera leo) on a moonless night, Masai Mara, Kenya. Taken with infra red camera, September
Snow Geese (Chen caerulescens) in flight, silhouetted against colourful dusk sky. Bosque del Apache, New Mexico, USA, November
Snow Goose (Chen caerulescens), coming in to land. Bosque del Apache, New Mexico, USA, November
Marine Iguana (Amblyrhynchus cristatus) on rocks with wave splash. Galapagos, Ecuador
Red Squirrel (Sciurus vulgaris) adult reaching up. Cairngorms National Park, Scotland, UK, March
Heermanns Gulls (Larus heermanni) in flight. Ensenada, Baja California Peninsula, Mexico, December
Monarch butterfly (Danaus plexippus) hibernating, Mariposa Monarca Special Biosphere Reserve, central Mexico, January
American Flamingo (Phoenicopterus ruber), Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, August
Polar bear (Ursus maritimus) adult walking over newly formed pack ice during fall freeze up, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska, USA
Polar bear (Ursus maritimus) sow and cub sliding on their backs over the pack ice, much like ice skating, 1002 coastal plain of the Arctic National Wildlife Refuge, Alaska, USA
Dall sheep (Ovis dalli) solitary animal at sunset with a storm moving across the sky, Denali National Park, interior Alaska, USA
Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA
Long Horn cattle in meadow, Norfolk, UK July
Ash (Fraxinus excelsior) fresh leaf in spring, Dorset, UK April
A school of lesser devil rays (Mobula hypostoma) flying through sunbeams as they feed. Isla Mujeres, Quintana Roo, Yucatan Peninsula, Mexico. Caribbean Sea
Whitetip reef sharks (Triaenodon obesus) pack silhouetted following scent trail in water column, Cocos Island, Costa Rica, Pacific Ocean
Sunset on Everest (8848m), Sagarmatha National Park (World Heritage UNESCO). Khumbu / Everest Region, Nepal, Himalaya, October 2011
Tajun Pokhari Lake (4. 600 m) with mountains behind, Sagarmatha National Park (WorldTajun Pokhari Lake (4.600 m) with mountains behind, Sagarmatha National Park (World Heritage UNESCO). Khumbu / Everest Region, Nepal, Himalaya, October 2011
Emperor angelfish (Pomacanthus imperator) with soft corals and sweepers. Egypt, Red Sea
Mt Fitz Roy and Laguna Los Tres, panoramic view, Fitzroy National Park, Argentina
Perito Moreno Glacier, panoramic view, Argentina, January 2010
Spectacled caiman (Caiman yacare) walking on a sandy shore at the edge of a river at night, Pantanal, Brazil
Little egret (Egretta garzetta) attempting to catch fish, Camargue, France, July
White terns (Gygis alba) flock in flight overhead, Christmas Island / Kiritimati, Pacific Ocean, July
Looking out across sea from deserted beach, Christmas Island / Kiritimati, July 2008
Killer WHale / Orca (Orcinus orca) bull and cow in Resurrection Bay, Kenai Fjords National Park, outside Seward, Alaska, May
Great Egrets (Ardea alba) territorial fight above nest colony, Everglades National Park, Florida, USA, February
Aerial view over subtropical mangrove wetlands of the Everglades National Park. Florida, USA, February 2012
Whooper swan (Cygnus cygnus) in snowfall. Martin Mere Wetlands Trust, Lancashire, UK, January. Animal Portraits category, British Wildlife Photographer Awards (BWPA) competition 2012
White bellied woodstar (Chaetocercus mulsant) three hovering around red flowers, Guango private reserve, Papallacta Valley, Andean Cloud Forest, East slope, Ecuador
Purple gallinule (Porphyrio martinicus) moving across waterlily covered surface, Everglades National Park, Florida, USA, March
Big fin reef squid (Sepioteuthis lessoniana) descends from open water to the reef at night. Arafur Sea, Triton Bay, West Papua, Indonesia
Whiskered auklet (Aethia pygmaea), Iony Island / Jonas Island, Sea of Okhotsk, Far East Russia, July
Poppies flowering in a field, near Norcia, Umbria, Italy. May 2005
Flock of Greater flamingo (Phoenicopterus ruber) in flight, Camargue, Southern France, November 2006
