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Great crested grebe (Podiceps cristatus) feeding feather to one of its chicks, Valkenhorst nature reserve, Valkenswaard, The Netherlands. August
Male Leach's sea star (Leiaster leachi) broadcast spawning, releasing streams of sperm from its arms. Kagoshima Prefecture, Kyushu, Japan
Mimika bobtail squid (Euprymna morsei), Hokkaido, Japan, Pacific Ocean
Blackbird (Turdus merula) and falling snow. Leicestershire, UK, February
Banded demoiselle dragonfly (Calopteryx splendens) male and female silhouetted, resting on reeds. Lower Tamar Lakes, Cornwall, England, UK. June
English oak tree (Quercus robur) with tyre swing. Monmouthshire, Wales, UK, winter
Swallowtail butterfly (Papilio machaon), wings open, resting on plant stem. The Netherlands. August
Marine iguana (Amblyrhynchus cristatus) and Sally-lightfoot crabs (Grapsus grapsus) on rocks at high-tide with sea spray, Galapagos Islands, South America
A female kingfisher (Alcedo atthis) fishing through an ice hole in winter. Leeds, Yorkshire, UK. January. Sequence
Male Black grouse (Tetrao / Lyrurus tetrix) displaying at lek. Kuusamo, Finland, April
Panoramic landscape of Welsh Borders, countryside at dawn, Monmouthshire, Wales, UK, Panorama, October
Wren (Troglodytes troglodytes) singing during the morning at St Aidens RSPB reserve near Leeds, UK. April
Black and white photo of a silhouette of a California sea lion (Zalophus californianus) pup in an underwater cavern with sun beams. Los Islotes, La Paz, Baja California Sur, Mexico
Weasel (Mustela erminea) in white winter coat standing upright in snow, Upper Bavaria, Germany, Europe. February
Ocelot (Leopardus pardalis) high up in tree, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Pallas's cat (Otocolobus manul) kitten waiting at den for mother to return, Mongolia. June. 2017. Filmed for BBC series Big Cats
Lightning strike over South Africa's largest coal processing plant. Sasol, Mpumalanga, South Africa, November, 2016
Female scalefin anthias (Pseudanthias squamipinnis) swimming in front of a school of glassfish (Apogonidae), whose shiny bodies are rendered into out of focus bokeh patterns. Laamu Atoll, Maldives
Trees silhouetted against erupting Cumbre Vieja volcano at night, La Palma, Canary Islands, September 2021
Tawny owls (Strix aluco) peering out from tree hollow, Dumfries and Galloway, Scotland, UK, May, 2021
Crested tit (Lophophanes cristatus), Sierra de Grazalema Natural Park, Southern Spain. September
Close up lava from volcanic eruption, Cumbre Vieja Volcano, La Palma, Canary Islands. September 2021
Female Red fox (Vulpes vulpes) walking along tree trunk in heavy fog at night. Light source behind the subject creates dramatic volumetric lighting. Vertes Mountains, Hungary
Mute swan (Cygnus olor) in mist, Bolam Lake Country Park, Northumberland, UK, November
Adult humpback whale (Megaptera novaeangliae) diving in deep water channel. Great Bear Rainforest, British Columbia, Canada. COP26 Countdown Photo Competition 2021 Finalist. (digitally stitched image)
Spawning Leaf corals (Pavona decussata). Spawning is synchronized over a large area with many coral colonies, Kyushu, Japan, Pacific Ocean. July
Female Northern pygmy squid (Idiosepius paradoxus) preparing to lay another egg on a blade of eelgrass (Zostera marina), Yamaguchi Prefecture, Japan
Pair of Japanese dwarf flying squirrels (Pteromys volans orii) during the reproductive season, on branch nlooking down, Hokkaido, Japan. The female is on the left, male right
Juvenile Mute swan (Cygnus olor) in early morning light, Valkenhorst Nature Reserve, The Netherlands, Europe. August
Robin (Erithacus rubecula) carrying a red berry in its beak as it takes off, Parainen Uto, Finland. October
Caribbean flamingo (Phoenicopterus ruber) chick, standing by its parent's legs in its nest, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June. Bookplate
Crescent-tail bigeye (Priacanthus hamrur) school on a coral reef, shot with a long exposure. North Male Atoll, Maldives. Indian Ocean
European robin (Erithacus rubecula), Reddish Vale Country Park, Greater Manchester, UK. January
Water droplets on leaf creating a natural pattern, Tresco Tropical Garden, Tresco, Isle of Scilly, Cornwall, UK. June
Banded demoiselle (Calopteryx splendens) male silhouetted on a fern, Lower Tamar Lakes, Cornwall, UK. July
Greenfinch (Carduelis chloris) and Goldfinch (Carduelis carduelis) fighting in snow, Poland. February
Willow Tit (Parus montanus) in snow, Mainua, Kajaani, Finland, March
Snow covered trees in the Sotos del Alfaro Natural Reserve, Alfaro, La Rioja province, Spain. January
Mute swan (Cygnus olor) in early morning light. Valkenhorst nature reserve, Valkenswaard, The Netherlands. May
Northern gannets (Morus bassanus) pair displaying amongst campion, Bempton Cliffs, Yorkshire, UK, May
Crested tit (Lophophanes cristatus) Sierra de Grazalema Natural Park. Southern Spain. August
Volcanic eruption with silhouette of tree, Cumbre Vieja Volcano, La Palma, Canary Islands. September 2021
Roe deer (Capreolus capreolus) feeding in field with flowering Poppies (Papaver rhoeas) and Cornflower (Centaurea cyanus), De Inslag, Brasschaat, Belgium, June
Oak tree (Quercus robur) growing among bracken (Pteridium aquilinum), autumn colour, The New Forest, Hampshire, UK. November 2016
Beech trees (Fagus sylvatica) and English bluebells (Hyacinthoides non-scripta) soft focus, West Wood, near Marlborough, Wiltshire, UK. May 2021
Luskentyre beach/sands, marram grasses and early morning sunlight, Isle of Harris, Outer Hebrides, Scotland, UK. October 2018
High tide and metal hand railings at Bude Sea Pool, Bude, North Cornwall, UK. May 2021
Common spotted orchids (Dactylorhiza fuchsii), Kingcombe Meadows, Dorset, UK, June
