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Starling murmuration, hundreds of thousands of starlings (Sturnus vulgaris) gather in the skies near their night time roost. Gretna Green, Scotland, January
Schiehallion reflected in Loch Rannoch at dawn, Perthshire, Scotland, UK. May 2017
Dawn / sunrise over the Segama River, with mist hanging over lowland Dipterocarp rainforestDawn/sunrise over the Segama River, with mist hanging over lowland Dipterocarp rainforest. Heart of Danum Valley, Sabah, Borneo
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Wren (Troglodytes troglodytes) singing, early morning in spring, Broxwater, Cornwall, UK. April
Resplendent quetzal (Pharomachrus mocinno) male, Talamanca mountains, Costa Rica
Male Bengal Tiger - Sundar (B2) (Panthera tigris tigris) running and patrolling territory. Bandhavgarh National Park, Madhya Pradesh, India
Saturniid moth (Aurivillius triramis), Laraganga Mole National Park, Ghana, Africa
Brown four eyed opossum (Metachirus nudicauda) at night in cloud forest, Choco region, Northwestern Ecuador. Camera trap image
Arctic fox cub (Alopex lagopus) portrait in grass with summer flowers, Hornvik, Hornstrandir, Westfjords, Iceland. July
Adolescent male Bengal Tiger (Panthera tigris tigris) (around 15 months) climbing a tree. Bandhavgarh NP, Madhya Pradesh, India
Autumn colour along meandering river bank, River Khovd, Altai Mountains, Bayan-Ulgii, Western Mongolia
Four Little bee-eaters (Merops pusillus) perched on branch, Seronera Valley, Serenegeti National Park, Tanzania
Sumatran (Asian) two-horned Rhinoceros (Dicerorhinus sumatrensis) found in isolated rainforest in Borneo and Sumatra. In captivity at Sepilok Rehabilitation Centre, Sabah, Borneo
Female ring-tailed lemurs (Lemur catta) carrying infants (3-4 weeks) on their backs across open ground. Berenty Private Reserve, southern Madagascar
Female Verreauxs sifaka (Propithecus verreauxi) carrying infant in forest canopy. Berenty Private Reserve, southern Madagascar
Giant Otters (Pteronura brasiliensis) running along a sand bar. Cuiaba River, Pantanal, Brazil
Eurasian jay (Garrulus glandarius) foraging in autumn leaves. London, England, UK. November
Male common blue butterfly (Polyommatus icarus) basking wings open on grass, Vealand Farm, Devon, UK. June
Pacific white-sided dolphins (Lagenorhynchus obliquidens) underwater, Queen Charlotte Strait, British Columbia, Canada. September
Giant Pacific octopus (Enteroctopus dofleini) showing arms and suckers, Vernon Rock, Queen Charlotte Strait, British Columbia, Canada. September
Black Grouse (Lyrurus tetrix) male displaying with bokeh effect in the foreground, Liminka, Finland, March
Portrait of two wild Bobcat (Lynx rufus) kittens in a tree, Texas, USA. September
Bairds tapir (Tapirus bairdii) walking along a beach in Corcovado National Park, Costa Rica. Endangered
Common blue damselfly (Enallagma cyathigerum) portrait, early morning light, Broxwater, Cornwall, UK. June 2018
Two hard fern (Blechnum spicant) fronds uncurling, Coombe Valley, Cornwall, UK. May
Slime mould (Physarum psittacinum), in mature reproductive phase. Close-up of erupting fruiting bodies (sporangia), bearing thousands of spores. Buckinghamshire, UK
Evening Grosbeak (Coccothraustes vespertinus) male perched on spruce branch with falling snow, in winter, New York, USA
A vibrant Caribbean coral reef with two Reef sharks (Carcharhinus perezi) and Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Monte Iberia eleuth (Eleutherodactylus iberia), on human fingernail, one of the Worlds smallest frogs. Endemic to Cuba. Humboldt National Park, Cuba
Male Korean seahorse (Hippocampus haema) giving birth. Kumamoto Prefecture, Kyushu, Japan
Bowhead whale (Balaena mysticetus) tail slapping, Sea of Okhotsk, Russia
Common octopus (Octopus vulgaris) hunting on a reef off Eleuthera Island, Bahamas
Panamic Cushion Sea Star (Pentaceraster cumingi) and a school of Yellow-tailed Grunt (Anisotremus interruptus), Galapagos Island, Ecuador. December
Caribbean reef squid (Sepioteuthis sepioidea) portrait reflecting off the surface of the ocean at night, Eleuthera, Bahamas
Hongotlyn Dunes, Gobi Desert, Mongolia. October 2008
Mute swan (cygnus olor) in the first light in the mist Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, June
Landscape with beautiful clouds on a windless day Corcovado National Park, Osa peninsula, Costa Rica
Arctic fox (Vulpes lagopus) cub in meadow, portrait. Hornstrandir Nature Reserve, Iceland, July
Portrait of Bornean Horned Frog (Megophrys nasuta) amongst leaf-litter in forest floor. Taken at night. Danum Valley, Sabah, Borneo
Male Panther Chameleon (Furcifer pardalis) stalking prey in beach side vegetation. Bay of Antongil, Masoala Peninsula National Park, north east Madagascar
Golden-crowned or Tattersalls sifaka (Propithecus tattersalli), forests near the village of Andranotsimaty, near Daraina, northern Madagascar
Black-browed Albatross (Thalassarche melanophrys) flying off the Bay of Isles, South Atlantic. January. (digitally stitched image)
King penguins (Aptenodytes patagonicus) on the beach at Salisbury Plain, with Antarctic Tern (Sterna vittata) hovering. South Georgia, South Atlantic
Buttermere reflections, Cumbria, The Lake District, UK. November 2016
Blackchurch Rock at Mouth Mill, near Browhsham / Clovelly, last light, North Devon, UK. April 2019
Westward Ho! beach at sunrise, colourful sky at low tide and tidal pool, north Devon, UK. February