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Mute swan (Cygnus olor) pair at sunrise. London, UK. December
Mute swan (Cygnus olor) cygnets and parents swimming. London, UK. May
Three Marbled white butterflies (Melanargia galathea) nectaring on Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June
Gyre flyingfish (Prognichthys glaphyrae) photographed in the Sargasso Sea, Atlantic Ocean
Ring tailed lemur (Lemur catta) pair on rocky outcrop, Anja Community Reserve, Ambalavao, Madagascar
Kaleidoscopic image of autumn leaves. UK
Kaleidoscopic montage of a peacock feather
Red-throated Diver (Gavia stellata) calling, Iceland, June
Red-throated Diver (Gavia stellata) taking off with fish prey, Iceland, June
Hazel Grouse (Tetrastes bonasia), Kuusamo, Finland, December
Black-throated Diver (Gavia arctica) Vaala Finland, June
White-tailed Eagle (Haliaetus albicilla) Norway, November
Mountains, Nubra Valley, Karakorum Wildlife Sanctuary, Ladakh, India. September 2011
Dead Langs short-tailed blue butterfly (Leptotes pirithous) in a river with bacterial film, Los Alcornocales Natural Park, Southern Spain, September
Common redstart (Phoenicurus phoenicurus) taking off, Sierra de Grazalema Natural Park. Southern Spain. September
Translucent coral goby (Bryaninops erythrops) in Seapen (Virgularia gustaviana). Komodo National Park, Indonesia. Indo-West Pacific
St Michael's Mount at sunrise with a rainbow over Penzance, viewed from Marazion, West Cornwall, UK. February 2019
Tigress (Panthera tigris tigris) with her four cubs from Chandrapur district, Maharashtra, India. She lives in an area known for human-animal conflict
Autumn crocus or meadow saffron (Colchicum autumnalis) in Lower Woods, Gloucestershire, England, UK, September
Honey garlic (Nectaroscordium siculum) with Bumblebee in flight. Naturalised in the Avon Gorge, Bristol, UK, June
Common snowdrops (Galanthis nivalis) growing in the Rococo Gardens, Painswick, Gloucestershire, Engalnd, UK. February
Monarch butterfly (Danaus plexippus) wintering in Oyamel pine forests (Abies religiosa) Monarch Butterfly Biosphere Reserve / Reserva de Biosfera de la Mariposa Monarca UNESCO World Heritage Site
Northern fulmar (Fulmarus glacialis) in flight with waves, Arctic Sea. Commended in the Biophoto Competition 2019
African elephant (Loxodonta africana) two silhouetted at sunset with goose flying overhead, Mkuze, South Africa. Highly commended in the African Wildlife category of the Nature's Best Photography
Supermoon rising over Stonehenge Wiltshire, the biggest Supermoon in 68 years and the closest since 1948. 13th November 2016
Californian sea lion (Zalophus californianus) swimming in Southern sea palm (Eisenia arborea) and Giant Kelp (Macrocystis pyrifera) forest, San Benitos Islands
Meadow grasshopper (Pseudochorthippus parallelus) on Common heather (Calluna vulgaris), covered in dew droplets. Klein Schietveld, Brasschaat, Belgium. August
Small emerald spreadwing (Lestes virens) pair mating. Klein Schietveld, Brasschaat, Belgium. August
Sun shining through tree at dawn. Klein Schietveld, Brasschaat, Belgium. July 2019
American oak (Quercus rubra) trees in avenue, sun shining through. Peerdsbos, Brasschaat, Belgium. November 2013
Goat willow (Salix caprea) in mist at dawn, full moon in sky. Klein Schietveld, Brasschaat, Belgium. August 2019
North American river otter (Lutra canadiensis) two on frozen river edge. Upper Yellowstone River, Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Greater flamingo (Phoenicopterus roseus) flock, Pont Du Gau Park, Camargue, France
Green sea turtle (Chelonia mydas), two viewed from above. Tenerife, Canary Islands
Arctic fox (Alopex lagopus) in snow. Svalbard, Norway. April
Reindeer (Rangifer tarandus), two play fighting in snow. Svalbard, Norway, April
Polar bear (Ursus maritimus) male female pair looking upwards, snow covered hills in background. Svalbard, Norway, April 2018
Alcedo giant tortoise (Chelonoidis vandenburghi), group bathing. Alcedo Volcano, Isabela Island, Galapagos. 2017
Southern elephant seal (Mirounga leonina), two males, equally matched with mouths open in aggression. Out of focus King penguin (Aptenodytes patagonicus) colony in background
Red fox (Vulpes vulpes) sequence of snow diving whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February 2019. Digital composite
Black-browed albatross (Thalassarche melanophris) in flight over sea at sunrise. South Atlantic Ocean between The Falklands and South Georgia. November
North American river otter (Lutra canadiensis), two standing at edge of frozen Upper Yellowstone River, Hayden Valley, Yellowstone, USA. February
Guanacos (Lama guanicoe), five silhouetted on slope in evening. Torres del Paine National Park, Patagonia, Chile. December
Fragrant orchids (Gymnadenia conopsea) amongst wildflowers in ancient alpine meadow. North Tyrol, Austria. June
King penguin (Aptenodytes patagonicus), two sleeping on snow. Grytviken, South Georgia. March
Spinner dolphin (Stenella longirostris) pod socialising close to surface. Fury Shoal, Sataya Reef, Marsa Alam, Egypt
Oceanic whitetip shark (Carcharhinus longimanus) swimming close to surface. Brothers Islands, Egypt
Snow leopard (Unica unica) standing beside boulder in rocky landscape of Himalayas, prior to scent marking. Ule, Ladakh, India. March 2018. Sequence 1/2
