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Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016
Red squirrel (Sciurus vulgaris), two curled up asleep in drey inside nest box. Nest of lichen and Pine needles. Highlands, Scotland, UK. Medium repro only
Snowdrop (Galanthus Sp. ) single flower in snow, Buckinghamshire, England, UK, FebruarySnowdrop (Galanthus Sp.) single flower in snow, Buckinghamshire, England, UK, February
Dugong (Dugong dugon) male feeding on a seagrass meadow (Halophila stipulacea), accompanied by a young Golden trevally (Gnathanodon speciosus). Marsa Nabaa, Marsa Alam, Egypt. Red Sea
Tarpon (Megalops atlanticus) portrait, Eleuthera, Bahamas. December
Polar bear (Ursus maritimus) standing on ice. Svalbard, Norway, July
Kiang (Equus kiang) herd grazing on steppe in evening light, mountains in background. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October 2019
House Martin (Delichon urbica) Spain, April
Eastern tiger salamander (Ambystoma tigrinum) North Florida, USA. December
Canadian timber wolf / Northwestern wolf (Canis lupus occidentalis) pack moving through snow, Wood Buffalo National Park, Alberta, Canada. Taken on location for BBC Frozen Planet, March 2009
Tre Cime di Lavaredo / Drei Zinnen, three distinctive mountain peaks in the Sexten Dolomites seen from WW1 cave shelter, South Tyrol, Italy, October 2019
Golden eagle (Aquila chrysaetus), Kuusamo, Finland, January
English bluebell (Hyacinthoides non-scripta) Lower Woods, Gloucestershire, England, UK, April
Elephant (Loxodonta africana) herd walking in a line, Etosha National Park, Namibia
White-tailed eagle (Haliaeetus albicilla) taking off with fish prey, Norway, August
Buckholt Wood is a beech (Fagus sylvatica) wood and part of the Cotwolds Commons and Beechwoods National Nature Reserve, on the scarp slope of the Cotswolds near Birdlip, Gloucestershire
Goshawk (Accipiter gentilis) plucking a Willow Grouse (Lagopus lagopus) Liminka, Finland, March
Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) crossing the Mara River. Northern Serengeti, Serengeti National Park, Tanzania
Grey wolf running in snow (Canis lupus), Minnesota, USA. January. Controlled situation
Lady Amhersts pheasant (Chrysolophus amherstiae) Kanding, Sichuan, China
Fosa of Fossa (Cryptoprocta ferox) male walking through dry deciduous forest, Kirindy, western Madagascar
Barn and wheat field in the Palouse farming area of southeastern Washington, USA, August
Golden eagle (Aquila chrysaetos) Sodankyla, Lokka, Finland, April
Sea elephant (Cardiopoda placenta), marine pelagic gastropod mollusc in the family Pterotracheidae. Deep sea heteropod. Atlantic Ocean, close to Cape Verde. Controlled conditions
Deadly nightshade (Atropa belladonna) Quarry Hangers (SWT), Chaldon, Surrey, England, June
Portrait of a wild male Bobcat (Lynx rufus), Texas, USA. September
Red fox (Vulpes vulpes) cubs sleeping curled up together, Switzerland
Central bearded dragon (Pogona vitticeps) male on stick in chenopod shrubland habitat, Meringur in NW Victoria, Australia
Autumn quaking aspen trees (Populus tremuloides) below Turret Ridge, Uncompahgre National Forest, Colorado, USA. October
White-tailed deer (Odocoileus virginianus) doe standing in snow, New York, USA
Common octopus (Octopus vulgaris) on a coral reef, The Bahamas. August
Golden snub-nosed monkey (Rhinopithecus roxellana), mother and baby, Qinling Mountains, Shaanxi province, China
Lion-tailed macaque (Macaca silenus), dominant male, Anaimalai Mountain Range (Nilgiri hills), Tamil Nadu, India
Southern cassowary (Casuarius casuarius) Bali Bird Park, Denpasar, Bali, Indonesia. Captive
Strawberry land hermit crab (Coenobita perlatus). On white coral sand beach, Small unhinhabited coral islet in Raa Atoll lagoon, Maldives
Clouded leopard (Neofelis nebulosa) portrait, captive, occurs in the Himalayas
Face portrait of a Great hammerhead shark (Sphyrna mokarran) off Bimini, Bahamas
Magpie Goose (Anseranas semipalmata) at sunrise Lotusbird Billabong, Cape York Peninsula Queensland, Australia
Eclectus parrot (Eclectus roratus) two males arrive at a nest cavity at the same time, both to provision the female within. Iron Range National Park, Cape York Peninsula, Queensland, Australia
Pallass cat (Otocolobus manul) licking lips whilst running over ice. East Mongolia. February
Umbrella thorn tree (Vachellia tortilis) in the Etosha pan with common ostrich (Struthio camelus) male. Etosha National Park, Namibia
Aerial view of Glassy Point, East Coast of Dominica, West Indies. December 2019. Drone photo. Stitched panorama
Hyacinth macaws (Anodorhynchus hyacinthinus) squabbling on palm tree, Pantanal, Mato Grosso, Brazil
Lion (Panthera leo) males mock fighting / play fighting. Ngorongoro Crater, Ngorongoro Conservation Area (NCA), Tanzania
Red fox (Vulpes vulpes) running through snow. Hayden Valley, Yellowstone, USA. January
Fosa / Fossa (Cryptoprocta ferox) male resting in dry decidous forest. Kirindy, western Madagascar. Endangered
Firefly (Lamprohiza splendidula) light trails of males in forest, Bavaria, Germany. July
Southern right whale (Eubalaena australis) diving, with tail fluke splash. Monumento Natural Ballena Franca Austral, UNESCO World Heritage Site, Valdes Peninsula, Patagonia, Argentina
Common Loons (Gavia immer) one diving underwater, Michigan, USA. June
Grey wolf in snow (Canis lupus), Minnesota, USA. January. Controlled situation
Red deer (Cervus elaphus) silhoutted at dusk, Richmond Park, London, England, UK. October
Moose bull (Alces alces) walking in forest clearing, Denali National Park, Alaska, USA, September
Dramatic sunset with storm clouds over Roseau, Caribbean sea view in Dominica, Lesser Antiles. September 2019
Great Grey Owl (Strix nebulosa) Kuhmo Finland, March
Gannet (Morus bassanus) Shetland, England, UK. July
Black Woodpecker male (Dryocopus martius) Finland, January. One branch removed
Sparrowhawk (Accipiter nisus) adult female, Norway, October
Golden jackal (Canis aureus) pup sitting on a grassy verge by a path, Danube Delta, Romania, July
Lady Amhersts pheasant (Chrysolophus amherstiae) male displaying, Kanding, Sichuan, China. April
European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition
Silhouette of an African leopard (Panthera pardus pardus) snarling, Mkuze, South Africa. September. White Fang - this leopard bared its fangs to the hyenas threatening his prey
Polar bears (Ursus maritimus) fighting in water. Beaufort Sea, Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Natures Best Photography Competition 2019
Cuckoo (Cuculus canorus) in flight, Germany, April. May
Common Cuckoo (Cuculus canorus) perched on gate Surrey, England, UK. April
Starling (Sturnus vulgaris) murmuration at sunset, West Pier, Brighton, England, UK
Aye-aye (Daubentonia madagascariensis) adult active and foraging in forest canopy at night. Deciduous forests, Daraina, northern Madagascar. Endangered endemic species
Red fox (Vulpes vulpes) adult hunting for rodents by snow diving in deep snow. Hayden Valley, Yellowstone, USA. January
Fosa or Fossa (Cryptoprocta ferox)(sometimes erroneously fossa) male in dry deciduous forest. Kirindy, western Madagascar
Puma (Puma concolor puma), female sitting in grassland. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. November
Puma (Puma concolor puma), young male crossing road in front of car. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December 2018
Alpine bearberry (Arctous alpina) on tundra in autumn and boulderrs at Sydkapp (South Cape), with icebergs in Hall Bredning, Scoresby Sund, Greenland, September
Giant panda (Ailuropoda melanoleuca) cub in tree, Chengdu Panda Breeding Centre, Sichuan, China. Captive
Fangtooth moray (Enchelycore anatina) with open mouth, portrait. Tenerife, Canary Islands
Ocean sunfish (Mola mola) and Pilot fish (Naucrates sp). Tenerife, Canary Islands
Napoleon wrasse (Cheilinus undulatus) on a coral reef. Ras Mohammed National Park, Sinai, Egypt. Red Sea
Bluestreak cleaner wrasse (Labroides dimidiatus) cleans among the sharp teeth of a predatory roving coral grouper (Plectropomus pessuliferus) beneath an overhang on a coral reef
Arctic fox (Alopex lagopus) camouflaged in winter pelage. Svalbard, Norway, April
Polar bear (Ursus maritimus) walking across ice, snow covered mountain in background. Svalbard, Norway, April 2018
Red fox (Vulpes vulpes) in mid air, snow diving / pouncing whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February
Puma (Puma concolor puma) young male standing with mouth open. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December
Guanaco (Lama guanicoe) female, alert whilst looking at Puma (Puma concolor) on on opposite slope, portrait. Torres del Paine National Park, Patagonia, Chile. November
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) swimming over Seagrass (Halophila stipulacea) meadow. Marsa Nabaa, Marsa Alam, Egypt
Aerial image of Luskentyre Beach on the Isle of Harris, Scotland, UK, August
Reef manta (Mobula alfredi) visiting a cleaning station on a coral reef with a school of Onespot snappers (Lutjanus monostigma)
