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African lion (Panthera leo) and lioness after mating, Masai Mara National Reserve, Kenya
Lion (Panthera leo) male walking. Masai Mara National Reserve, Kenya
Polar bears (Ursus maritimus) fighting in water. Beaufort Sea, Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Natures Best Photography Competition 2019
Fosa of Fossa (Cryptoprocta ferox) male walking through dry deciduous forest, Kirindy, western Madagascar
Red fox (Vulpes vulpes) adult hunting for rodents by snow diving in deep snow. Hayden Valley, Yellowstone, USA. January
Boky-boky (Mungotictis decemlineata) or Narrow-striped mongoose adult foraging in leaf litter. Dry deciduous forest, Kirindy, western Madagascar
Fosa or Fossa (Cryptoprocta ferox)(sometimes erroneously fossa) male in dry deciduous forest. Kirindy, western Madagascar
Puma (Puma concolor puma), female sitting in grassland. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. November
RF- Wild male Jaguar (Panthera onca palustris) running through shallows of backwater of Cuiaba River in late afternoon sunlight. Northern Pantanal, Brazil
Giant River Otter (Pteronura brasiliensis) feeding on Striped Catfish or Cachara (Pseudoplatystoma fasciatum) amongst water hyacinth. Tributary off the Cuiaba River, Mato Grosso, Pantanal, Brazil
Atlantic walrus (Odobenus rosmarus rosmarus) hauled out on ice floe, Svalbard, Norway June
Giant panda (Ailuropoda melanoleuca) cub in tree, Chengdu Panda Breeding Centre, Sichuan, China. Captive
Arctic fox (Alopex lagopus) camouflaged in winter pelage. Svalbard, Norway, April
Polar bear (Ursus maritimus) standing on ice. Svalbard, Norway, July
Lion (Panthera leo) drinking at night, Zimanga private game reserve, KwaZulu-Natal, South Africa
Red fox (Vulpes vulpes) in mid air, snow diving / pouncing whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February
Guanaco (Lama guanicoe) female, alert whilst looking at Puma (Puma concolor) on on opposite slope, portrait. Torres del Paine National Park, Patagonia, Chile. November
Siberian weasel (Mustela sibirica) standing in snow. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
Giant river otter (Pteronura brasiliensis) resting on fallen tree trunk above Cuiaba River. Pantanal, Mato Grosso, Brazil
Melanistic leopard / Black panther (Panthera pardus fusca) resting in tree. Nagarhole National Park, India. Photo© Phillip Ross/Felis Images
Fishing cat (Prionailurus viverrinus) walking with fish in mouth. Andhra Pradesh, India. Camera trap image. Photo© Anjani Kumar/Felis Images
Asiatic lion (Panthera leo persica), two females and two cubs. Gir National Park, Gujarat, India. Photo© Phillip Ross/Felis Images
Red fox (Vulpes vulpes) dog interacting with a vixen in an urban garden. North London, UK. June
Giant river otter (Pteronura brasiliensis) eating a catfish. Yasuni National Park, Orellana, Ecuador
Jungle cat (Felis chaus) at night, Kanha National Park and Tiger Reserve, Madhya Pradesh, India
Galapagos sea lion (Zalophus wollebaeki) juvenile on beach, Galapagos
A female American black bear (Ursus americanus) and her young cub, Nova Scotia, Canada. July
Bengal tiger (Panthera tigris tigris) (T27) walking on bund / dam wall trail. Kanha National Park, Central India. Camera trap image
Tiger (Panthera tigris tigris) dominant male tiger approaching Beja pani waterhole, Kanha National Park, Central India. Camera trap image
Young grey seal pup (Halichoerus grypus) recently born on a beach in Orkney, Scotland, UK, April
Red Fox (Vulpes vulpes) running through deep snow. London, UK. January
Stoat (Mustela erminea) Creux du Van, Jura Suisse, Switzerland, September
Badger (Meles meles), cub pulling the ear of sibling, June, Scotland, UK. August
Fox cub (Vulpes vulpes) in late summer. Dorset, UK, August
Common Seals (Phoca vitulina) hauled out on sand bank at Blakeney Point, Norfolk, England, UK, August
Tayra (Eira barbara) climbing in a tree in rainforest habitat with fern covered trees, Bellavista private reserve, Mindo cloud forest area, Ecuador, July
Red Fox (Vulpes Vulpes) next to red fox graffiti art at night. North London, England UK
Giant panda (Ailuropoda melanoleuca) sitting with Bamboo in background. Shenshuping Panda Base, Wolong Nature Reserve, Wenchuan, Sichuan Province, China. Captive
Spotted necked otters (Hydrictis maculicollis), Chobe River, Botswana, September
European otter (Lutra lutra) two at waters edge, one shaking off water, Kiskunsagi National Park, Pusztaszer, Hungary, October
European otters (Lutra lutra) feeding at water?s edge with splashing water, Kiskunsagi National Park, Hungary, January
RF- North American river otter (Lutra canadensis) captive, occurs in North America
Giant Otter (Pteronura brasiliensis) holding onto a branch in a lagoon off the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Oriental small clawed otter (Aonyx cinerea) underwater, captive
Asian / Oriental short-clawed otter (Aonyx cinerea) looking out of water with mouth open, Captive, Vulnerable species
Colonies of King penguins 1+Aptenodytes patagonicus+1 and Southern Elephant Seals 1+Mirounga leonina+1 on beach at St. Andrews Bay, South Georgia. November
Asian black bear (Ursus thibetanus) feeding on leaves, Tangjiahe Nature Reserve, Sichuan Province, China, April
Tibetan wolf (Canis lupus chanco) in mountain landscape, Serxu County, Garze Prefecture, Sichuan Province, China