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German Shepherd Dog bitch pup, Coco, 14 weeks old, with raised paw
American Red Fox (Vulpes vulpes) cubs playing with its mother. Grand Teton National Park, USA, June
Wild Cat (Felis silvestris) in mist. Vosges, France, August
Lion cub (Panthera leo) silhouetted at sunrise, taken on location for Pride tv series. September 2003
Blue and white Border Collie puppy running forward
Grey wolf (Canis lupus) mother and two month old cubs, pups begging for food by licking mothers mouth, captive
Grey wolves (Canis lupus) in snow, captive
Mexican wolf (Canis lupus baileyi), Mexican subspecies, probably extinct in the wild, captive
Close-up portrait of a European grey wolf (Canis lupus), captive, Norway, February
Two European grey wolves (Canis lupus), captive, Norway, February
European wolf (Canis lupis) in boreal birch forest in winter, Nord-Trondelag, Norway. Captive release
Arctic Fox (Alopex / Vulpes lagopus) at sunset, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Grizzly bear (Ursus arctos horribilis) cub peering out from behind its mother, Khutzeymateen Grizzly Bear Sanctuary, British Columbia, Canada, June
Male Arabian Leopard (Panthera pardus nimr) at the Arabian Wildlife Centre & captive-breeding project, Sharjah, United Arab Emirates
On a misty morning, female Grizzly bear (Ursus arctos horribilis) standing up in alert and looking around with her cubs for danger, while feeding on sedges, Khutzeymateen Grizzly Bear Sanctuary
Arctic fox (Alopex lagopus) Svalbard, Norway, April
Brown bear (Ursus arctos) and cubs in the snow, Finland, April
RF - Polar Bear (Ursus maritimus) males fighting, Churchill, Canada, November
Grizzly Bears (Ursus arctos) resting, Lake Clarke National Park, Alaska, September
Grizzly Bear (Ursus arctos) asleep, Lake Clarke National Park, Alaska, September
Brown Bear (Ursus arctos) adult portait, Finland, June
Black bears (Ursus americanus) three cubs, one climbing, Minnesota, USA, June
Coastal brown bear (Ursus arctos) female in a meadow, cubs nearby, Lake Clarke National Park, Alaska, USA, July
Polar bear (Ursus maritimus) curious young male on the newly frozen pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Polar bear (Ursus maritimus) 3-year-olds play fighting on newly formed pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Pine marten (Martes martes) leaping, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Animal Behaviour category of the BWPA (British Wildlife Photography Awards) Competition 2016
Lion (Panthera Leo) male, low angle shot, South Africa. Taken on location for Pride tv series
Female Sea lion (Zalophus wollebaeki) with pup suckling in shallow water, Floreana Island, Galapagos
Galapagos sea lion (Zalophus wollebaeki) pup on beach, Floreana Island, Galapagos, Endangered species
Sri Lankan leopard (Panthera pardus kotiya) Yala National Park, Southern Province, Sri Lanka
North-western wolf (Canis lupus occidentalis) portrait, captive occurs in northwestern USA and Canada
Lioness (Panthera leo) chasing away Cheetah (Acinonyx jubatus) Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Badger (Meles meles) adult in pine forest at sunset, Derbyshire, UK, June. Highly commended in the Habitat category of the British Wildlife Photography Awards (BWPA) Competition 2017
Common otter (Lutra lutra) Norfolk UK May
European badger (Meles meles) foraging in deciduous woodland. June, Mid Devon, UK
Young Red fox (Vulpes vulpes) portrait, Kent, UK, June
Badger (Meles meles) at entrance to sett, Derbyshire, England, UK, June. Highly commended in the Habitat category of the BWPA competition 2014
Red Fox (Vulpes vulpes) portrait in an urban area. Glasgow, Scotland. May
Steller sealion (Eumetopias jubatus) male charging past the camera. Race Rocks, Victoria, Vancouver Island, British Columbia. Canada. North East Pacific Ocean
California sea lions (Zalophus californianus) group playing in the sun in the early morning. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Polar bear (Ursus maritimus) silhouetted at waters edge at sunset, Bernard Spit, off the 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, September. Vulnerable species
Indochinese tiger (Panthera tigris corbetti) looking up, behind screen of leaves, captive, occurs in South East Asia
Young Polar bear (Ursus maritimus) walking across melting sea ice, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
Red fox (Vulpes vulpes) silhouetted at dusk, after sunset, The Netherlands, July
Leopard Seal (Hydrurga leptonyx) resting on iceberg, Antarctica, February
Cheetah (Acinonyx jubatus) resting on a termite hill and storm brewing, Masai-Mara Game Reserve, Kenya. March
Cheetah (Acinonyx jubatus) female with cubs sitting in long grass during the dry season, Masai-Mara Game Reserve, Kenya. March
Cheetah (Acinonyx jubatus) female standing below dark storm clouds, Masai-Mara game reserve, Kenya
