Black Bear (Ursus americanus) sleeping in a Sitka Spruce tree (Picea sitchensis) alongside Anan Creek, Alaska, July
Brown bear (Ursus arctos arctos) female and two cubs, aged 1 year, emerging from den in winter, Bavarian Forest, Germany, captive, January
African wild dogs (Lycaon pictus) juveniles playing with the leg of an impala, trying to drag it in three different directions. Mkuze, South Africa
Polar bear (Ursus maritimus) walking on Champ Island glacier above the sea, Franz Josef Land, Russian Arctic. Highly commended in Animals in their Environment Category of the Wildlife Photographer of
Arctic Fox (Vulpes lagopus), in winter coat portrait, Svalbard, Norway, April
Snow leopard (Panthera uncia) portrait with ears back. Captive
Grizzly bear (Ursus arctos horribilis) female with cub riding on back, Katmai National Park, Alaska, USA, August
Killer whale (Orcinus orca) male named Mel attacking young South American sea lion (Otaria flavescens) on beach, Punta Norte, Peninsula Valdes, Argentina
Red fox (Vulpes vulpes) vixen in front of Clifton Suspension Bridge at night. Avon Gorge, Bristol, UK. March 2016
Young urban Red fox (Vulpes vulpes) poking its head up over a wall. Bristol, UK, August
Red fox (Vulpes vulpes) cubs playing, Vosges, France, May
RF - Lion (Panthera leo) head portrait at night, Zimanga private game reserve, KwaZulu-Natal, South Africa. (This image may be licensed either as rights managed or royalty free.)
Wild Jaguar (Panthera onca), Endangered, Cuiaba River, Pantanal, Mato Grosso, Brazil
RF - Lions (Panthera leo) - two brothers patrolling territorial boundary. Short grass plains on the border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Polar bear (Ursus maritimus) young bear rolling around in the snow, on newly formed pack ice during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Young urban Red fox (Vulpes vulpes) standing on a wall at night. Bristol, UK, September. Nominated in the Melvita Nature Images Awards competition 2014
Urban Red fox (Vulpes vulpes) walking past wall with red fox mural / graffiti. North London, England, UK, April
African lion (Panthera leo) portrait, Masai Mara Game Reserve, Kenya
Grey seal (Halichoerus grypus) swimming towards camera, Orkney, Scotland, UK, August
Sea otter (Enhydra Lutris) with forelegs raised, Sitka Sound, Alaska, USA, August
Polar bear cubs (Ursus maritimus) triplets age 2-3 months next to their mother. Wapusk National Park, Manitoba, Canada. February
Arctic fox (Vulpes lagopus), juvenile looking at camera, portrait, winter pelage. Dovrefjell National Park, Norway. September
Wild female Amur leopard (Panthera pardus orientalis) on rocky hillside, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species
Grey Wolf (Canis lupus), in forest, captive, USA
Giant Panda cub (Ailuropoda melanoleuca) climbing. Yuan Meng, first Giant panda even born in FranceGiant Panda cub (Ailuropoda melanoleuca) climbing.Yuan Meng, first Giant panda even born in France, is now aged 8 months. and likes very much to stay in trees, Beauval Zoo, France
Pine marten (Martes martes) portrait, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Portrait category of the BWPA (British Wildlife Photography Awards) Competition 2016
RF - Arctic Fox (Vulpes lagopus) portrait in winter coat, Svalbard, Norway, April
Giant panda (Ailuropoda melanoleuca) cub yawning. Yuan Meng, first giant panda ever born in France, age 10 months, Captive at Beauval Zoo, Saint Aignan sur Cher, France
Male Fossa (Cryptoprocta ferox) climbing down tree trunk from forest canopy. Mid-altitude rainforest, Andasibe-Mantadia National Park, eastern Madagascar. Endangered
Arctic fox (Vulpes lagopus), juvenile running through snow, winter pelage. Dovrefjell National Park, Norway. September
Lions (Panthera leo) - two brothers patrolling territorial boundary, affectionate behaviour, border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Giant panda cub (Ailuropoda melanoleuca) portrait Yuan Meng, first giant panda ever born in France, age 10 months. Captive at Beauval Zoo, Saint Aignan sur Cher, France
Brown bear (Ursus arctos) male, portrait. Martinselkonen, Kainuu, Finland. June
Sable (Martes zibellina) in tree, Irkutsk, Siberia, Russia. November
Red fox (Vulpes vulpes) in winter snow, Jura, Switzerland
Canadian timber wolf / Northwestern wolf (Canis lupus occidentalis) pack moving through snow, Wood Buffalo National Park, Alberta, Canada. Taken on location for BBC Frozen Planet, March 2009
Bengal Tiger (Panthera tigris) six month old cub jumping on its mother, Bandhavgarh National Park, India
Arctic fox (Vulpes lagopus) portrait of juvenile, winter pelage. Dovrefjell National Park, Norway. September
Common eider duck (Somateria mollissima) female defending her nest from arctic fox (Vulpes lagopus). Longyearbyen, Spitsbergen. June
Arctic fox (Vulpes lagopus) juvenile sniffing ground, winter pelage. Dovrefjell National Park, Norway. September
Snow leopard (Panthera uncia) in snow, captive, USA
Leopard (Panthera pardus) female fighting off male after he tries to mate with her. Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Polar bear (Ursus maritimus) on an iceberg, Baffin Bay, Canada. September
RF - Snow leopard (Panthera uncia) female, portrait, captive
Snow leopard (Uncia uncia) Altai Mountains, Mongolia. March
Grizzly bear (Ursus arctos) catching a fish, Brooks Falls in Katmai National Park, Alaska, USA, July
Jaguar (Panthera onca) male walking, Pantanal, Brazil
Grey wolf (Canis lupus) adult greeted by cub, captive, USA