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Pine marten (Martes martes) reflected in water, Ardnamurchan Peninsula, west coast of Scotland, UK. Highly commended in the Mammals category of the British Wildlife Photography Awards (BWPA)
Weasel (Mustela nivalis) looking out of hole on woodland floor with snowdrops, Sheffield, England, UK
Young urban Red fox (Vulpes vulpes). Bristol, UK. August
Siberian tiger (Panthera tigris altaica) in snow, captive
Cougar (Puma concolor) portrait, captive, occurs in Americas. Digitally manipulated
RF - Lion (Panthera leo) male on savanna with dramatic storm clouds, thundery sky and light rays in the background. Ngorongoro Crater, Tanzania. Composite image
Bengal Tiger (Panthera tigris) six month old cub jumping on its mother, Bandhavgarh National Park, India
Meerkats (Suricata suricatta) with young, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
African wild dog (Lycaon pictus) interacting with pup age two months, Okavango Delta, Botswana
African Wild dogs or Cape hunting dogs (Lycaon pictus) at close range taken from ground level, Zimanga Private Game Reserve, South Africa
Snow leopard (Panthera uncia) in snow, captive, USA
Bobcat (Lynx rufus) standing on branch in snow. Madison River Valley, Yellowstone National Park, Wyoming, USA. January
Brown bear (Ursus arctos) paw seem from under water, Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August
Cheetah (Acinonyx jubatus) staring back over its shoulder, black and white. Save Valley Conservancy, Zimbabwe, August
Red panda (Ailurus fulgens) captive, occurs in China
Timber Wolves (Canis lupus) resting on a rock by the shoreline. Near Mussel Inlet, Great Bear Rainforest, British Columbia, Canada, October
Pine marten (Martes martes) climbing Scots pine tree, Scotland, UK. May
Lioness (Panthera leo) carrying her cub, Masai-Mara Game Reserve, Kenya
Grey wolf (Canis lupus) adult greeted by cub, captive, USA
Pine marten (Martes martes) portrait, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Portrait category of the BWPA (British Wildlife Photography Awards) Competition 2016
Badger (Meles meles) adult, backlit at sunset, scratching its ears, Derbyshire, UK, May. Winner of the Animal Behaviour category of the British Wildlife Photography Awards (BWPA) Competition 2017
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Black Bear (Ursus americanus) sleeping in a Sitka Spruce tree (Picea sitchensis) alongside Anan Creek, Alaska, July
Giant panda (Ailuropoda melanoleuca) cub yawning. Yuan Meng, first giant panda ever born in France, age 10 months, Captive at Beauval Zoo, Saint Aignan sur Cher, France
Polar bear cub 3 months (Ursus maritimus) playing in the front of the day den in march. Wapusk National Park. Churchill, Manitoba, Canada
Lion (Panthera leo) cubs playing, Masai-Mara Game Reserve, Kenya. Vulnerable species
Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek against ancient Manchurian fir tree, Land of the Leopard National Park, Far East Russia, November
Red fox (Vulpes vulpes) in snowy landscape with trees and two Crows (Corfus corone) winter, Jura, Switzerland
Sumatran tiger (Panthera tigris sondaica). Captive, with digitally added leaf pattern
Coyote (Canis latrans) walking through deep winter snow. Yellowstone National Park, Wyoming, USA. January
Wild Eurasian lynx (Lynx lynx) at night with city lights and sky glow behind, Switzerland. December. Taken with remote camera trap
Brown bear (Ursus arctos) portrait in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia
Kamchatcka brown bear (Ursus arctos beringianus) with face underwater whilst fishing for Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Russia, August
Badger (Meles meles) cub amongst long grass, Dorset, England, UK, July
Red fox (Vulpes vulpes) resting in the snow, Churchill, Cananda, November
Australian sea (Neophoca cinerea) juvenile playing in the sand, Carnac Island, Western Australia
Australian sea lion (Neophoca cinerea) playful juvenile swimming upside-down, Carnac Island, Western Australia
Brown bear (Ursus arctos arctos) female and two cubs, aged 1 year, emerging from den in winter, Bavarian Forest, Germany, captive, January
Grey Seal (Halichoerus grypus) at rest on submerged rock, head and one flipper above water. Shetland Islands, Scotland, UK. July
Grey wolf running in snow (Canis lupus), Minnesota, USA. January. Controlled situation
Arctic fox (Alopex lagopus) in the snow, in summer coat, Svalbard, Norway, April
Arctic Fox (Vulpes lagopus) portrait in winter coat, Svalbard, Norway, April
Portrait of a meerkat (Suricata suricatta) or suricate, looking at the camera and standing to survey its territory on the edge of Makgadikgadi Pans National Park, Botswana, April
Polar bears (Ursus maritimus) standing up on hind legs, barrier island outside Kaktovik, Alaska, USA. September
Arctic fox (Alopex lagopus), in winter coat portrait, Svalbard, Norway, April
Asian leopard cat (Prionailurus bengalensis) captive, occurs in South East Asia
Domestic dog, Weimaraner, puppy in garden, France
Arctic wolf (Canis lupus) male and female, captive