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Brown bear (Ursus arctos) and Grey wolf (Canis lupus) together in wetlands, Kuhmo, Finland, July
African wild dogs (Lycaon pictus) juveniles playing with the leg of an impala, trying to drag it in three different directions. Mkuze, South Africa
Golden Cocker Spaniel puppy, Maizy, sitting
Arctic Fox (Vulpes lagopus), in winter coat portrait, Svalbard, Norway, April
Red fox (Vulpes vulpes) vixen in front of Clifton Suspension Bridge at night. Avon Gorge, Bristol, UK. March 2016
Young urban Red fox (Vulpes vulpes) poking its head up over a wall. Bristol, UK, August
African Wild Dog (Lycaon pictus) pack passing infront of large African elephant (Loxodonta africana) Mana Pools National Park, Zimbabwe October 2012
Red fox (Vulpes vulpes) cubs playing, Vosges, France, May
Young urban Red fox (Vulpes vulpes) standing on a wall at night. Bristol, UK, September. Nominated in the Melvita Nature Images Awards competition 2014
Urban Red fox (Vulpes vulpes) walking past wall with red fox mural / graffiti. North London, England, UK, April
Arctic fox (Vulpes lagopus), juvenile looking at camera, portrait, winter pelage. Dovrefjell National Park, Norway. September
Golden Retriever puppy in basket with flowers. USA
Grey Wolf (Canis lupus), in forest, captive, USA
Cavalier King Charles Spaniel, puppy, black-and-tan, 6 weeks, sitting on grass, wearing collar
RF - Arctic Fox (Vulpes lagopus) portrait in winter coat, Svalbard, Norway, April
Young Red fox (Vulpes vulpes) in garden outside Royal Fort House, Bristol University, UK, January
Red fox (Vulpes vulpes) in winter snow, Jura, Switzerland
Canadian timber wolf / Northwestern wolf (Canis lupus occidentalis) pack moving through snow, Wood Buffalo National Park, Alberta, Canada. Taken on location for BBC Frozen Planet, March 2009
Arctic fox (Vulpes lagopus) portrait of juvenile, winter pelage. Dovrefjell National Park, Norway. September
Common eider duck (Somateria mollissima) female defending her nest from arctic fox (Vulpes lagopus). Longyearbyen, Spitsbergen. June
Arctic fox (Vulpes lagopus), juvenile running through snow, winter pelage. Dovrefjell National Park, Norway. September
Arctic fox (Vulpes lagopus) juvenile sniffing ground, winter pelage. Dovrefjell National Park, Norway. September
Red fox (Vulpes vulpes) at night in snow, camera trap image, Jura Mountains, Switzerland, August. Commended in the Mammals category of the Asferico Competition 2016
Red fox (Vulpes vulpes) portrait in snow, Kamchatka, Far east Russia, April
Arctic Fox (Alopex / Vulpes lagopus) standing on ridge, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Arctic Fox (Alopex / Vulpes lagopus) yawning, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Young urban Red fox (Vulpes vulpes). Bristol, UK. August
African wild dog (Lycaon pictus) interacting with pup age two months, Okavango Delta, Botswana
African Wild dogs or Cape hunting dogs (Lycaon pictus) at close range taken from ground level, Zimanga Private Game Reserve, South Africa
Timber Wolves (Canis lupus) resting on a rock by the shoreline. Near Mussel Inlet, Great Bear Rainforest, British Columbia, Canada, October
Grey wolf (Canis lupus) adult greeted by cub, captive, USA
Red Fox (Vulpes vulpes) cub in late evening light, Leicestershire, England, UK, July
Wolf (Canis lupus) pack running in deep snow, captive in enclosure of the Bavarian Forest National Park, Germany, February
Red fox (Vulpes vulpes) in snowy landscape with trees and two Crows (Corfus corone) winter, Jura, Switzerland
Coyote (Canis latrans) walking through deep winter snow. Yellowstone National Park, Wyoming, USA. January
Red fox (Vulpes vulpes) resting in the snow, Churchill, Cananda, November
Grey wolf running in snow (Canis lupus), Minnesota, USA. January. Controlled situation
Arctic fox (Alopex lagopus) in the snow, in summer coat, Svalbard, Norway, April
Arctic Fox (Vulpes lagopus) portrait in winter coat, Svalbard, Norway, April
Two Dogue de Bordeaux dogs looking over garden fence. No release available
Arctic fox (Alopex lagopus), in winter coat portrait, Svalbard, Norway, April
German Shepherd / Alsatian, puppy, 4 months, sitting with tongue out
Domestic dog, Weimaraner, puppy in garden, France
Arctic wolf (Canis lupus) male and female, captive
RF- European grey wolf (Canis lupus) running through snow in birch forest, Tromso, Norway. Captive, April
Arctic fox (Alopex lagopus) young fox pouncing, Iceland. October
Arctic fox (Vulpes lagopus) walking through snowy landscape, Taymyr Peninsula, Siberia, Russia
Red fox (Vulpes vulpes) silhouetted in mixed beech-Douglas fir forest. Freiburg, Black Forest National Park, Germany, October