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Orca whales (Orcinus orca) pod surfacing together, Shetland, Scotland, UK. April
Caribou / Reindeer bull (Rangifer tarandus) close-up grazing, Denali National Park, Alaska, USA, September
Moose (Alces alces) bull swimming in water, Baxter State Park, Maine, USA
Rocky mountain elk (Cervus elaphus nelsoni) bull bugling during rut. Rocky Mountain National Park, Colorado, USA. October
Southern elephant seal (Mirounga leonina) beach master / bull roaring amongst females and pups, in breeding colony. St Andrews Bay, South Georgia. October
Orca (Orcinus orca) surfacing and blowing, Shetland, Scotland, UK, August
Muskox (Ovibos moschatus), male portrait. Dovrefjell National Park, Norway
African elephant (Loxodonta africana) bull with dried mud covering skin. Mud creates a monochrome texture. Savuti, Botswana. Taken on location for BBC Planet Earth series, 2005
Matador waving red cape at bull during bullfight, bull is speared with barbed sticks (banderillas), Plaza de Toros, Valencia, Spain. July 2014
Moose (Alces alces) bull crossing mountain river at sunset, Grand Teton National Park, Wyoming, USA
Moose (Alces alces) bull, portrait. Grand Teton National Park, Wyoming, USA. September
Moose (Alces alces) bull swimming across river. Grand Teton National Park, Wyoming, USA. September
Moose (Alces alces) bull, crossing mountain river after sunset. Grand Teton National Park, Wyoming, USA. October 2017
Orca (Orcinus orca) surfacing, Shetland, Scotland, UK, August
Moose (Alces americanus) close-up grazing, Denali National Park, Alaska, USA, September
Sable antelope (Hippotragus niger) bull charging through the shallows of the Selinda Spillway, with another behind, northern Botswana, August
Male Black rhinoceros (Diceros bicornis), Phinda Private Game Reserve, Kwazulu Natal, South Africa, February
Nilgai (Boselaphus tragocamelus) young bulls, Keoladeo / Bharatpur National Park UNESCO Natural World Heritage Site, India
Camargue bull (Bos taurus) running through marshland, Camargue, France. October
Bull Moose (Alces alces) drinking from mountain stream, Grand Teton National Park, Wyoming, USA. October
Moose (Alces alces) two bulls sparring in willow flat grasslands, Grand Teton National Park, Wyoming, USA. October
African elephant (Loxodonta africa) male in threat display to rival. Floor of the Ngorongoro Crater, Ngorongoro Conservation Area, Tanzania
Cow with cowbell silhouetted at sunrise, with mist over the Eastern Rhodope Mountains, Madzharovo, Bulgaria, November 2014
Grey seal (Halichoerus grypus) on haul out in fishing harbour with ferry in the background, Shetland Isles, Scotland, UK, June 2010
Grey seal (Halichoerus grypus) bull with reflections on water of harbour lights, Shetland Isles, Scotland, UK, June 2010. Photographer quote
Herd of Camargue bulls, Camargue, France, May
Moose Bull (Alces alces) walking in forest clearing, Denali National Park, USA, September
Black rhino (Diceros bicornis) and White rhino (Ceratotherium simum), being harrassed by playfighting White rhino bull, Solio Game Reserve, Laikipia, Kenya. September
