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Blue-footed booby (Sula nebouxii) courting pair, South coast, Santa Cruz Island, Galapagos
Blue-footed booby (Sula nebouxii) landing, South coast, Santa Cruz Island, Galapagos
Great frigatebird (Fregata minor) in tree with pair of Nazca boobies (Sula granti) showing mating behaviour, Genovesa Island, Galapagos
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
Two Red-footed booby (Sula sula) perched on branches, Genovesa Island, Galapagos Islands, Ecuador
Male Blue-footed booby (Sula nebouxii) walking on sand, San Cristobal Island, Galapagos, South America
Red-footed booby (Sula sula), adult and chick at nest. Genovesa Island, Galapagos
Nazca booby (Sula granti) preening. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Blue-footed booby (Sula nebouxii) landing, Santa Cruz Island, Galapagos, Ecuador
Red-footed booby (Sula sula), pair looking at one another on nest in tree. Egg is nest. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair perched in tree. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair nest building in tree. Darwin Bay, Genovesa Island, Galapagos
Nazca booby (Sula granti) at nest with egg. Overlooking sea, Gardner Islet, Floreana Island, Galapagos. December 2014
Nazca booby (Sula granti), pair amongst Prickly pear (Opuntia sp) cacti at coast. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Blue-footed booby (Sula nebouxii), six diving into sea. Espumilla Beach, Santiago Island, Galapagos
Red-footed booby (Sula sula), brown morph, Genovesa Island, Galapagos Islands
Nazca booby (Sula granti) on the coast of Wolf Island, Galapagos
Magnificent frigatebird (Fregata magnificens) attacking Blue footed booby (Sula nebouxii) feeding chick. This bird is trying to steal the food from the parent. Santa Cruz Island, Galapagos
Blue-footed booby (Sula nebouxii) with chicks begging for food, Santa Cruz Island, Galapagos, Ecuador
Red-footed booby (Sula sula) perched in tree. Genovesa Island, Galapagos
Nazca booby (Sula granti), pair on rocky shore at dawn. Punta Suarez, Espanola Island, Galapagos. May 2015
Nazca booby (Sula granti) standing on rock at coast. Punta Suarez, Espanola Island, Galapagos. October 2015
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
Nazca booby (Sula granti) portrait, Galapagos, Ecuador
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Brown booby (Sula leucogaster), adult diving for fish, Los Roques, Venezuela
Red Footed Booby (Sula sula) couple, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
Red-footed booby (Sula sula) perched on Palo santo tree (Bursera graveolens), with flowering Yellow cordia (Cordia lutea) in background. Galapagos Islands, Ecuador
Red-footed booby (Sula sula) chick and parent perched on branch near nest, Genovesa Island, Galapagos Islands, Ecuador
Red-footed booby (Sula sula) perched on branch, Genovesa Island, Galapagos Islands, Ecuador
Great frigatebird (Fregata minor) attempting to steal food from Nazca booby (Sula granti) feeding its chick, Genovesa Island, Galapagos Islands, Ecuador
Blue-footed booby (Sula nebouxii), incubating eggs with webbed feet, Seymour Island, Galapagos, South America
Blue-footed booby (Sula nebouxii) pair with chicks, blowhole in background, Punta Suarez, Espanola Island, Galapagos Islands
Brown booby (Sula leucogaster), Los Islotes, Espiritu Santo Archipelago National Park, Sea of Cortez (Gulf of California), Mexico, September
Blue-footed booby (Sula nebouxii), diving, Santa Cruz Island, Galapagos Islands
Red-footed booby (Sula sula) landing among salt bushes, looking for nesting materials, with Frigate birds behind, Genovesa Island, Galapagos
Nazca booby (Sula granti) on coast, with Vampire ground finches (Geospiza septentrionalis). These finches peck at birds, feeding on blood when other food sources are scarce, Wolf sland, Galapagos
Nazca booby (Sula granti) on nest. Genovesa Island, Galapagos. February 2017
Nazca booby (Sula granti) in flight at dusk. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) on sea, tossing fish. Turtle Cove, Santa Cruz Island, Galapagos
Blue-footed booby (Sula nebouxii) plunge-diving at high speed, San Cristobal Island, Galapagos, Ecuador
Booby (Sula nebouxii) looking at camera underwater, Galapagos Island
Brown booby (Sula leucogaster) flying over San Pedro Martir Island Protected Area, Gulf of California (Sea of Cortez), Mexico, July