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Killer whale (Orcinus orca) hunting inside herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Yellow warty frogfish (Antennarius maculatus) Seraya, Tulamben area, Bali, Indonesia. Java Sea
Silver Drummer (Kyphosus sydneyanus) fish off the coast of Neds Beach, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Giant Manta Ray (Manta birostris), with Remoras (Echeneididae) Socorro Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico, March
Young Stone sculpin (Paracottus knerii), on endemic sponge (Lubomirskia baicalensis), Lake Baikal, Siberia, Russia
Tropical Flounder (Bothus mancus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
European Chubb (Squalius / Leuciscus cephalus) in cave, Plitvice Lakes National Park, Croatia. November
Orange anemonefish (Amphiprion sandaracinos) in its host Mertens carpet sea anemone (Stichodactyla mertensii). Mabul Island, Sabah, Borneo, Malaysia. Celebes Sea / Sulawesi Sea
Flame anthias (Pseudanthias ignitus) displaying to the more drab female (above) during courtship in the evening. Similan Islands, Thailand. Andaman Sea, Indian Ocean
Great hammerhead shark (Sphyrna mokarran) female at night with Bar jacks (Caranx ruber), South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Patagonian redfish (Sebastes oculatus) about 4cm, Comau Fjord, Patagonia, Chile
Mandarinfish (Synchiropus splendidus) Yap, Micronesia
Giant moray eel (Gymnothorax javanicus) opening its mouth to breathe, at night on the barge wreck, Gubal Island, northern Red Sea
Dwarf lionfish (Dendrochirus brachypterus) opens its awingsa (in fact enlarged pectoralDwarf lionfish (Dendrochirus brachypterus) opens its wings (in fact enlarged pectoral fins) Seraya, Tulamben area, Bali, Indonesia, Java Sea
Bignose unicornfish (Naso vlamingii) profile of male displaying (the blue stripe at the bottom of the frame is the tail of another individual) Baa Atoll, Maldives. Indian Ocean
Blackspotted stingray (Taeniurops meyeni) with a remora (Echeneis naucrates) attached to its belly swimming in the open water with a diver in the background. Ari Atoll, Maldives, Indian Ocean
Killer whale (Orcinus orca) adult male surfacing at dusk surrounded by birds, who watch for herrings (Clupea harengus) left after they have fed, Andenes, Andoya island, North Atlantic Ocean, Norway
Galapagos barnacle blenny (Acanthemblemaria castroi) looking out from its home in an old barnacle shell. Cape Marshall, Isabela Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Clarks anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia
Bigeye trevally (Caranx sexfasciatus) forming a circular shoal in blue water, on the drop off of a coral reef. Ras Mohammed National Park, Sinai, Egypt. Red Sea
Smooth trunkfish (Lactophrys triqueter), portrait. Bonaire, Dutch Caribbean
Black Bream (Spondyliosoma cantharus) washed up on shore, Sark, British Channel Islands, August
Short snouted seahorse juvenile (Hippocampus hippocampus) Sark, British Channel Islands, August
Ling (Molva molva) St Abbs Voluntary Marine Reserve, Scotland (North Sea)
Eggs of Long-spined sea scorpion (Taurulus bubalis) on sea shore, possibly being predated by Painted Topshell, Guernsey, British Channel Islands
RF - Orange skunk clownfish (Amphiprion sandaracinos) in anemone, Puerto Galera, Philippines
Big-eye or Goggle-eye (Priacanthus hamrur) and Crown squirrelfish (Sargocentron diadema) on wreck colonised by corals, Red Sea, Egypt. January
Kaleidoscopic image of coral reef with soft corals (Dendropnethya) shoal of Yellow back fusilliers (Caesio teres) West Papua, Indonesia
RF - Coral reef with Soft corals (Dendronephthya sp) and fish, West Papua, Indonesia
Spinecheek anemonefish (Premnas biaculeatus) male with bleached anemone, Bunaken National Park, North Sulawesi, Indonesia
Big-eye fish (Priacanthus hamrur). Egypt, Red Sea
Emperor angelfish (Pomacanthus imperator) swimming past coral reef. Egypt, Red Sea
Yellowbar angelfish (Pomacanthus maculosus) Egypt, Red Sea
Gorgonian sea fan (Melithaea sp. ) with Lyretail anthias (Pseudanthias sqamipinnis)Gorgonian sea fan (Melithaea sp.) with Lyretail anthias (Pseudanthias sqamipinnis). Similan Islands, Andaman Sea, Thailand
RF - Shoal of Slender pinjalo snappers (Pinjalo lewisi) swimming over soft corals on coral reef. West Papua, Indonesia
Ring-tailed Cardinalfish (Apogon aureus) West Papua, Indonesia
Crown spotty blenny (Laiphognathus longispinis) Pak Lap Tsai, Sai Kung East Country Park, Hong Kong, China
Blue shark (Prionace glauca) with Pilot fish (Naucrates ductor) off Halifax, Nova Scotia, Canada. July
Piraputanga (Brycon hilarii) and small Tetra fish (Hyphessobrycon sp. ) Rio Olhio d AguaPiraputanga (Brycon hilarii) and small Tetra fish (Hyphessobrycon sp.) Rio Olhio d Agua, tributary of Rio da Prata, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul
Atlantic salmon (Salmo salar) jumping up waterfall during spawning migration upstream, Ume River, Sweden. July. Photographed for The Freshwater Project
Sargassum fish (Histrio histrio) at the sea surface with floating sargassum weed. Hawaii
Painted cardinalfish (Archamia fucata) Yap, Micronesia
European barracuda (Sphyraena sphyraena) and Bluefish (Pomatomus saltatrix) circling baitball of Atlantic horse mackerel (Trachurus trachurus) Formigas Islets, Azores
Bullethead parrotfish (Chlorurus sordidus) male, close up of eye, whilst sleeping at night. The Alternatives, Sinai, Egypt. Red Sea
Whitebelly damselfish (Amblyglyphidodon leucogaster) pair spawning / laying eggs on a seafan (Annella mollis). Ras Mohammed National Park, Sinai, Egypt. Red Sea
Goby (Bryaninops sp) on coral branch, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Reef and shoal of Jewel fairy basslet (Pseudanthias squamipinnis) South Point dive site, Sanganeb reef, Sudan, Red Sea