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Ocean sunfish (Mola mola) and Pilot fish (Naucrates sp). Tenerife, Canary Islands
Napoleon wrasse (Cheilinus undulatus) on a coral reef. Ras Mohammed National Park, Sinai, Egypt. Red Sea
Bluestreak cleaner wrasse (Labroides dimidiatus) cleans among the sharp teeth of a predatory roving coral grouper (Plectropomus pessuliferus) beneath an overhang on a coral reef
Flying fish (Exocoetidae) in Sargasso Sea, Atlantic Ocean
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) swimming over Seagrass (Halophila stipulacea) meadow. Marsa Nabaa, Marsa Alam, Egypt
Chinese sturgeon (Acipenser sinensis), captive in aquarium. Adults are no longer able to swim upriver to spawn due to Gezhouba and Three Gorges dams
Bubble eye goldfish (Carassius auratus) with upward pointing eyes and two large fluid-filled sacs.. In captivity, Shanghai Zoo, Shanghai, China
Chinese grass carp / White amur (Ctenopharyngodon idella), farmed for food in China. Captive specimen, China
Blue shark (Prionace glauca) with a pair of pilot fish (Naucrates ductor) off Halifax, Nova Scotia, Canada. July
Moorish idol (Zanclus cornutus) juvenile with transparent skin in the open ocean at night off Anilao, Philippines
Pelagic thresher shark (Alopias pelagicus) being cleaned by a bluestreak cleaner wrasse (Labroides dimidiatus) at Monad Shoal off Malapascua Island, Philippines
Ribbon eel ( Rhinomuraena quaesita) Lembeh Strait, North Sulawesi, Indonesia
Taylors pygmy leatherjacket (Brachaluteres taylori). Lembeh Strait, North Sulawesi, Indonesia
Striped marlin (Kajikia audax) hunting Sardine (Sardinops sagax) in bait ball. Magdalena Bay, Baja California Sur, Mexico
School of Cow breams (Sarpa salpa ) Azores, Atlantic ocean
Trumpetfish (Aulostomus chinensis) on the coral drop off, Sulu Sea, Philippines
Invasive Lionfish (Pterois volitans) which have taken over and are wiping out native fish in the Atlantic ocean. The highest densities are in the northern gulf of Mexico. Destin, Florida, USA
Clownfish (Amphiprion sp) in anemone home, Philippines
Bigeye trevally (Caranx sexfasciatus), a large shoal. Cocos Island National Park, Costa Rica
Squirrelfish (Holocentrus sp) shoal amongst rocks. Near Cocos Island National Park, Costa Rica
Blue-lined snapper (Lutjanus kasmira) shoal. Cocos Island National Park, Costa Rica
Fish seeking shelter around floating tree in open ocean. Off coast of Cocos Island National Park, Costa Rica
Blacksmiths (Chromis punctipinnis) school swam through a giant kelp (Macrocystis pyrifera) forest, Santa Barbara Island, Channel Islands. Los Angeles, California, USA, North East Pacific Ocean
Nurse shark (Ginglymostoma cirratum) and Horse-eye jacks (Caranx latus) Shark Ray Alley, Hol Chan Marine Reserve, Belize Barrier Reef, Belize
Skeleton of a seahorse (Hippocampus sp. ) showing the remarkable structure of bonesSkeleton of a seahorse (Hippocampus sp.) showing the remarkable structure of bones that form a virtual exoskeleton, with few deep internal bones, in this unusual vertebrate group
Kingfisher (Alcedo atthis) male, after diving, taking off from water with fish, a Common Roach (Rutilus rutilus) Lorraine, France, August
Young European eel (Anguilla anguilla) elvers, or glass eels, caught during their annual migration up rivers from the Bristol channel, swimming in a large holding tank at UK Glass Eels
Giant Manta (Manta birostris) with Remoras (Remora brachyptera) attached to both wings, offshore Santa Maria Island, Azores, Atlantic Ocean
Wolf-eel (Anarrhichthys ocellatus) Vancouver Island, British Columbia, Pacific Ocean
Killer whale (Orcinus orca) diving and hunting for herring fish (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Great barracuda (Sphyraena barracuda) with three parasitic Copepods at end of upper jaw. Hawaii
Gorgonian / Sea whip coral (Ellisella ceratophyta) with Ring tailed cardinalfish (Apogon aureus) swimming in front, West Papua, Indonesia
Coral reef with soft corals (Dendronephthya sp. ), encrusting sponges and Pygmy sweepersCoral reef with soft corals (Dendronephthya sp.), encrusting sponges and Pygmy sweepers (Parapriacanthus ransonetti) Similan Islands, Andaman Sea, Thailand
Thread-tail dartgoby (Ptereleotris hanae) Sharp Island or Kiu Tsui Chau, Kiu Tsui Country Park, Hong Kong, China
School of Perch (Perca fluviatilis) amongst Water lilies (Nuphar lutea) lit by the suns rays, Lake Bourget, Alps, Savoie, France, June
Paddle-flap scorpionfish (Rhinopias eschmeyeri) Puerto Galera, Philippines
Devil scorpionfish (Scorpaenopsis diabolus) close up of eye, Pak Lap Tsai, Sai Kung, Hong Kong, China
Stellar rockskipper (Entomacrodus stellifer) hiding in a barnacle. Sai Kung East Country Park, Hong Kong, China
Ocean sunfish (Mola mola) off Halifax, Nova Scotia, Canada. July
Atlantic wolfish (Anarhichas lupus) looking out hole, Bay of Fundy, New Brunswick, Canada, July
Atlantic salmon (Salmo salar) portrait, migrating to spawn in river, Gaspe Peninsula, Quebec, Canada, October
Three spot damselfish (Dascyllus trimaculatus) with Sea anemone home, Yap, Micronesia
Common anemonefish (Amphiprion perideraion) with eggs in Magnificent sea anemone (Heteractis magnifica) Yap, Micronesia
Striped mackerel (Rastrelliger kanagurta) with mouths wide open as they swim through the water, filtering zoopankton with their gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Bluestreak cleaner wrasse (Labroides dimidiatus) sticking its head into the mouth of a Red Sea fusilier (Caesio lunaris) during a cleaning session. Small Crack, Sha ab Mahmud, Sinai, Egypt. Red Sea
Coral Rabbitfish (Siganus corallinus) portrait, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Cichlids (Cichlidae) in Lake Malawi, Malawi, November. Photographed for The Freshwater Project