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Atlantic Puffins (Fratercula arctica) pair bill rubbing, part of ritual courtship, Skomer Island, Wales, UK, May
Great crested grebes (Podiceps cristatus) performing courtship ritual at dawn, Cheshire, UK April
Female Giant panda (Ailuropoda melanoleuca), Huan Huan, holding her female cub, Yuandudu, age 3 months, Beauval Zoo, St.Aignan, France, November 2021. Captive
Spotted owlet (Athene brama) pair on a branch, Bharatpur / Keoladeo Ghana National Park, India
European rabbit (Oryctolagus cuniculus) two young rabbits, or kittens, touch noses, Norfolk, UK, June. Non-ex
Grizzly bear (Ursus arctos) female and nuzzling cub, Lake Clark National Park, Silver Salmon, Alaska. July
Infant Chimpanzee (Pan troglodytes troglodytes), aged 7 months, clinging onto its mother, Conkouati-Douli National Park, Republic of Congo, Africa
American oystercatchers (Haematopus palliatus) courting pair performing Piping Display, calling as they walk along together, Fort DeSoto Park, Florida, USA, March
Grizzly bear (Ursus arctos horribilis) cub peering out from behind its mother, Khutzeymateen Grizzly Bear Sanctuary, British Columbia, Canada, June
African lions (Panthera leo) one lioness licking another, Okavango Delta, Botswana
African Lion (Panthera leo) cub reaches for a moment of intimacy with its father, Northern Tuli Game Reserve, Botswana
Giraffe (Giraffa camelopardalis) female bending down to calf, Okavango Delta, Botswana
Grizzly bear (Ursus arctos) female with cub seeking attention from its mother, Lake Clark National Park, Silver Salmon, Alaska. July
Jaguar (Panthera onca) cub nuzzling its mother, Cuaiba River, Pantanal wetlands, Mato Grosso, Brazil
Two Cheetah (Acinonyx jubatus) brothers grooming one another. bonding behaviour, Linyanti, Chobe National Park, Botswana
Great crested grebe (Podiceps cristatus) carrying a chick on its back in early morning light, Valkenhorst Nature Reserve, The Netherlands, Europe. May
Two Lionesses (Panthera leo) nuzzling surrounded by three cubs, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Cheetah (Acinonyx jubatus) two leaning close together, Okavango Delta, Botswana
Pair of Gannets (Morus bassanus) mutual preening, Bass Rock, Firth of Forth, Scotland, UK, June
Great crested grebes (Podiceps cristatus) performing their elaborate courtship ritual at dawn, Cheshire, UK November
