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Harpy Eagle (Harpia harpyja) portrait. Gamboa, Soberania National Park, Panama
Female Harpy Eagle (Harpia harpyja) in flight. Returning to its nest. Pousada Currupira d Araras, south west Brazil
Bald eagle (Haliaeetus leucocephalus) pair flying with claws linked during courtship flight, Homer, Alaska, USA, March
Silver Birch (Betula pendula) trees in autumn colour, Craigellachie National Nature Reserve, Aviemore, Cairngorms National Park, Scotland, UK, October
Blue sky reflected in ripples of water, Svalbard, Norway, June 2010
Great frigatebird (Fregata minor) in tree with pair of Nazca boobies (Sula granti) showing mating behaviour, Genovesa Island, Galapagos
House Martin (Delichon urbica) Spain, April
Kestrel (Falco tinnunculus) male hovering overhead, Cornwall, UK, April
Oldshoremore Beach and dunes in evening light, Kinlochbervie, Sutherland, Scotland, UK, April 2014
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Lammergeier / Bearded vulture (Gypaetus barbatus) in flight with the out-of-focus moon behind. Ladakh, Himalayas, northern India
Three Red-crowned cranes (Grus japonensis) in flight. Hokkaido, Japan. March
The Angels Peak, Cairn Toul and Sgor an Lochan Uaine from Braeriach, Cairngorms National Park, Scotland, UK, September 2013
Central bearded dragon (Pogona vitticeps) male on stick in chenopod shrubland habitat, Meringur in NW Victoria, Australia
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Aerial image of Luskentyre Beach on the Isle of Harris, Scotland, UK, August
Nazca booby (Sula granti) preening. Punta Suarez, Espanola Island, Galapagos
RF- Waves of the North Sea washing onto Weybourne beach, Norfolk, UK. August 2014
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Red-collared lorikeet (Trichoglossus rubritorquis) pair examining hole in Eucalyptus as a potential nesting hollow. Katherine, Northern Territory, Australia
Common swift (Apus apus) group flying overhead, Bradford-on-Avon, Wiltshire, UK, May
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Irregular ripples on gypsum sand dunes created by high winds, White Sands National Monument, Chihuahuan Desert, New Mexico, USA, December 2012
Snowy egret (Egretta thula) portrait, viewed from below. St. Petersburg, Florida, USA
Pallass cat (Otocolobus manul) in snow, Gobi Desert, Mongolia. December
Northern pitcher plant (Sarracenia purpurea) photographed on Borgles Island, Nova Scotia, Canada, September
Pallass cat (Otocolobus manul) walking in snow, Gobi Desert, Mongolia. December
Gila monster (Heloderma suspectum) Arizona, USA. Captive
Indian Flying Fox (Pteropus giganteus), flying, India
Juvenile male Peregrine falcon (Falco peregrinus) in flight chasing his parent who has Feral pigeon (Columba livia) kill in claws, Bristol, England, UK, June
Goldfinch (Carduelis carduelis) perching on lichen covered branch, Glenfeshie, Scotland, UK, January
Waves lashing the Four Lighthouse during winter storm, Northern Brittany, France, December 2011. Sequence 8/12
Peregrine falcon (Falco peregrinus), adult male in flight. Bristol, UK. March
Alpaca Herd on Norfolk Farm, winter
Snowy owl (Bubo scandiacus) in flight, Boundary Bay, British Columbia, Canada. February. First place in the Overall Category of the 2013 Great Backyard Bird Count photo competition
Ash Tree (Fraxinus excelsior) growing in a field. Peak District National Park, Derbyshire, UK. August
Common tern (Sterna hirundo) in flight over water, Hampstead Heath, London, UK, August
Flock of Common Starling (Sturnus vulgaris) gathering before landing at winter roost, Salisbury Plain, Wiltshire, UK, January
RF - Coquerels sifaka (Propithecus coquereli) jumping, Anjajavy Private Reserve, north west Madagascar
Stars and Milky Way over Durdle Door and the Jurassic Coast, with the lights of Weymouth and Portland beyond, Dorset, England, UK, December 2013
Grants zebra (Equus burchelli boehmi) herd with large cloud above, Amboseli National Park, Kenya
Chalk rock formations caused by sand storms, White desert in the Sahara, Egypt, February 2009
European praying mantis (Mantis religiosa) camouflaged on leaf, Tuscany, Italy
White sand dunes against blue sky, White Sands National Monument, New Mexico, USA, February 2009
Tree covered with rime ice standing in snow-covered field, aginst blue sky with clouds, Ithaca, New York, USA. January 2010
Canadian Rockies reflected in marshland, Banff National Park, Alberta, Canada
White waterlily (Nymphaea alba) in Naardermeer bog lake, Holland. May
