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Bittern (Botaurus stellaris) in reeds, Suffolk, England, UK, November
King penguins 1+Aptenodytes patagonicus+1 group of three on the shore, St. Andrews Bay, South Georgia. November
King penguins 1+Aptenodytes patagonicus+1. Grytviken, South Georgia. November
Colonies of King penguins 1+Aptenodytes patagonicus+1 and Southern Elephant Seals 1+Mirounga leonina+1 on beach at St. Andrews Bay, South Georgia. November
Malachite kingfisher (Alcedo cristata) Bijagos Archipelago. Guineau Bissau
Osprey (Pandion haliaetus) fishing. Rothiemurchus, Cairngorms National Park, Scotland, UK
Rufous motmot (Baryphthengus martii) perched on branch. Rancho Naturalista, Cartago, Costa Rica
Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica
Violet sabrewing hummingbird (Campylopterus hemileucurus) nectaring on Orchid. Costa Rica
Red-legged partridge (Alectoris rufa) with chicks, Vendee, France, June
Little owl (Athene noctua) Sado Estuary, Portugal. June
White spoonbill (Platalea leucorodia) adults at nest with chicks, Sado estuary, Portugal. May
Common redstart (Phoenicurus phoenicurus) nest with six eggs, Alsace, France
Red-footed booby (Sula sula), adult and chick at nest. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair looking at one another on nest in tree. Egg is nest. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair perched in tree. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair nest building in tree. Darwin Bay, Genovesa Island, Galapagos
Nazca booby (Sula granti) at nest with egg. Overlooking sea, Gardner Islet, Floreana Island, Galapagos. December 2014
Nazca booby (Sula granti), pair amongst Prickly pear (Opuntia sp) cacti at coast. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Lapwing (Vanellus vanellus) portrait in upland hay meadow, Upper Teesdale, CO Durham, England, UK, June
Toco toucan (Ramphastos toco) acrobatically perched, feeding in forest canopy. Northern Pantanal, Cuiaba River, Mato Grosso, Brazil
Blue-footed booby (Sula nebouxii), six diving into sea. Espumilla Beach, Santiago Island, Galapagos
Trumpeter swans (Cygnus buccinator) on the edge of the Upper Yellowstone River. Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Burrowing owl (Athene cunicularia) doing rain dance, spreading wings and dancing while it rained, Arizona, USA
Cuckoo (Cuculus canorus) male, Germany, April
Blackcap (Sylviya atricapilla) male perched in blossom, Hungary, April
Red-collared lorikeet (Trichoglossus rubritorquis) pair examining hole in Eucalyptus as a potential nesting hollow. Katherine, Northern Territory, Australia
Great bowerbird (Chlamydera nuchalis), male showing off treasures whilst female inspects bower. Lake Argyle, Kununurra, Western Australia
Burrowing owl (Athene cunicularia) rotating head, Arizona, USA
Kestrel (Falco tinnunculus) fighting Hungary, April
Common Snipe (Gallinago gallinago) autumn courtship display, Germany, August
Blackbird (Turdus merula) in flight to feed on berries, Helsinki, Finland. November
Bearded reedling (Panurus biarmicus) on frosty reed seed head, Helsinki, Finland, January
Great grey owl (Strix nebulosa) taking off from snowy ground, Kuusamo, Finland. March
Capercaillie (Tetrao urogallus) male displaying in rain, Kuusamo, Finland. March
Red-legged honeycreeper (Cyanerpes cyaneus) two males on barbed wire. Costa Rica
Flock of Tree sparrows (Passer montanus) in flight, with a Greenfinch (Chloris chloris) blurred motion, Pacsmag, Hungary
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
Male Golden Pheasant (Chrysolophus pictus) standing on grass in display plumage. Kew Gardens, England, UK. March
Pine grosbeak (Pinicola enucleator), male feeding juvenile, Finland, July
Black grouse (Lyrurus tetrix), male at lek, Finland, April
Long-tailed duck (Clangula hyemalis), males in flight, Finland, April
Kingfisher (Alcedo atthis) male, after diving, taking off from water with fish, a Common Roach (Rutilus rutilus) Lorraine, France, August
Common swift (Apus apus) group flying overhead, Bradford-on-Avon, Wiltshire, UK, May
White-throated magpie-jay (Calocitta formosa) Nicoya peninsula, Costa Rica. February
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Eurasian hoopoe (Upupa epops) at nest in stone wall with hungry chicks gaping, Vendee, France, June