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Brown Roan Italian Spinone puppy, Riley, 13 weeks, chewing his tail
Little owl {Athene noctua) pair perched, courtship behaviour, Spain
African wild dogs (Lycaon pictus) juveniles playing with the leg of an impala, trying to drag it in three different directions. Mkuze, South Africa
Orangutan {Pongo pygmaeus} baby swinging in the trees, Rehabilitation sanctuary, Tanjung Puting National Park, Kalimantan, Indonesia
Little owls courtship {Athene noctua} Spain
Sperm whales (Physeter macrocephalus) resting, Pico, Azores, Portugal, June 2009 WWE OUTDOOR EXHIBITION. NOT AVAILABLE FOR GREETING CARDS OR CALENDARS. WWE BOOK
Chocolate roan Cocker Spaniel puppy, Topaz, 12 weeks, running with ears flapping
Barn owl (Tyto alba) in flight, hunting, Hampshire, England, UK. Captive
Dartford warbler (Sylvia undata) singing on Gorse, Dorset, UK, April
Blue-footed booby (Sula nebouxii) courting pair, South coast, Santa Cruz Island, Galapagos
Dugong (Dugong dugon) male feeding on a seagrass meadow (Halophila stipulacea), accompanied by a young Golden trevally (Gnathanodon speciosus). Marsa Nabaa, Marsa Alam, Egypt. Red Sea
Humpback whale (Megaptera novaeangliae) breaching, near Hout Bay, South Africa
Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October
Killer whale (Orcinus orca) male named Mel attacking young South American sea lion (Otaria flavescens) on beach, Punta Norte, Peninsula Valdes, Argentina
Robin (Erithacus rubecula) displaying in snow, nr Bradworthy, Devon, UK. December 2010, HIGHLY COMMENDED, ANIMAL PORTRAITS, 2011 WILDLIFE PHOTOGRAPHER OF THE YEAR COMPETITION
Hawksbill turtle (Eretmochelys imbricata) swimming underwater, Nosy Be, North Madagascar
Norwejian Fjord stallion running in the snow, Berthoud, Colorado, USA
Acorn Woodpecker (Melanerpes formicivorus), male at granary tree showing many acorns stored for winter survival, Orange County, California, USA
Male Pileated Woodpecker (Dryocopus pileatus) with beetle larva in beak about to feed two chicks, New York, USA, May
Leaf cutter ants (Atta sp) carrying colourful plant matter, reflected in water, Laguna del Lagarto, Santa Rita, Costa Rica
Bottlenose dolphin (Tursiops truncatus) playing in the waves of an oil tanker, Port Aransas, Texas, USA
Polar bear (Ursus maritimus) young bear rolling around in the snow, on newly formed pack ice during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
African Wild Dog (Lycaon pictus) pack passing infront of large African elephant (Loxodonta africana) Mana Pools National Park, Zimbabwe October 2012
Red fox (Vulpes vulpes) cubs playing, Vosges, France, May
Pipistrelle bat (Pipistrellus pipistrellus) flying low over water. Surrey, England, August
European grey wolf (Canis lupus) interacting with a European brown bear (Ursus arctos) Kuhmo, Finland, July 2009. WWE INDOOR EXHIBITION
Caribbean Flamingo (Phoenicopterus ruber) chick under the wing of protective parent, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico
Bottlenose dolphin (Tursiops truncatus) two breaching in evening light, Moray Firth, Inverness-shire, Scotland, UK, August, sequence 2/2
Snowy Owl (Nyctea scandiaca) adult female flying over snow, winter, Europe Captive /Snowy Owl (Nyctea scandiaca) adult female flying over snow, winter, Europe Captive/ Falconers bird, Austria
Adelie penguins (Pygoscelis adeliae) diving off iceberg, Antarctica, January
Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April
Dipper (Cinclus cinclus) singing from prominent rock in breeding season, Derbyshire, UK
Working English Springer Spaniel puppy, 6 weeks, in playful stance
Blacktail Prairie Dog (Cynomys ludovicianus) engaging in Jump-yip behavior - A strong arch of the back or " jump" followed by a shrill " yip" - Thought to occur when a predator
Common opossum {Didelphis marsupialis} playing dead as defence strategy
Platypus {Ornithorhynchus anatinus} swimming underwater Tasmania, Australia. Commposite
Mating pair of chalkhill blue butterflies (Lysandra coridon) with wings closed resting on Devils-bit scabious (Succisa pratensis), Hatch Hill, Somerset, UK. August
Bald eagle (Haliaeetus leucocephalus) pair flying with claws linked during courtship flight, Homer, Alaska, USA, March
Portrait of a Golden snub-nosed monkey (Rhinopithecus roxellana) screaming and showing its teeth, Foping Nature Reserve, Shaanxi, China. Endangered species
Adelie penguin (Pygoscelis Adeliae) in falling snow on the western Antarctic Peninsula, Southern Ocean
Killer whale (Orcinus orca) pod hunting together in herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Playful tortoiseshell kitten
Osprey (Pandion haliaetus) in flight catching a fish, Finland, July
Wild female Amur leopard (Panthera pardus orientalis) on rocky hillside, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species
Puffin (Fratercula arctica) swimming underwater, Farne Islands, Northumberland, UK, July
Golden Oriole (Oriolus oriolus) male feeding its chicks at nest, Bulgaria
Japanese crane (Grus japonensis) displaying, wings in heart shape, Hokkiado, Japan, February
White-vented storm petrel (Oceanites gracilis galapagoensis) running across water, offshore Cape Berkeley, Isabela island, Galapagos