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White ibis (Eudocimus albus) group of four in flight above clouds, Fort Myers Beach, Gulf Coast, Florida, USA, March
RF- Mute swan (Cygnus olor) on water in winter dawn mist. Loch Insh, Cairngorms National Park, Highlands, Scotland UK, December
Hen Harrier (Circus cyaneus) adult female in flight over heather moorland, Scotland, UK. July
Hoatzins (Opisthocomus hoazin) perched in tropical rainforest, Tambopata Reserve, Peru, South America
Tree sparrow (Passer montanus) feeding a fledgling on a branch of Hawthorn (Crataegus monogyna) blossom, Perthshire, Scotland. June 2013
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Yellow-plumed parakeets (Leptosittaca branickii) at nest cavity in wax palm stump, Tapichalaca Biological Reserve, Ecuador
Barn owl on post in flooded Somerset Levels. England, UK, Europe
Bateleur eagle (Terathopius ecaudatus) head portrait, Captive, occurs in Africa
Long-eared owl (Asio otus) in a large tree, Cheshire, UK, March
Wren (Troglodytes troglodytes) on frost covered fern, Dunham Massey, Cheshire, UK. Winner of the Habitat Category of the British Wildlife Photographer of the Year Awards (BWPA) 2017
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Adelie penguin (Pygoscelis adeliae) diving near ice flow, Antarctica. Small reproduction only
Himalayan monal (Lophophorus impejanus) male perched on branch, Uttarakhand, India
Golden eagle (Aquila chrysaetus) Kuusamo, Finland, January
Coal tit (Periparus ater) on a branch with lichen, Vendee, France, December
Golden plover (Pluvialis apricaria) chick camouflaged in ground nest, Norway June
Three Red-crowned cranes (Grus japonensis) in flight. Hokkaido, Japan. March
Common loon (Gavia immer), in breeding plumage, running across water to take flight, Michigan, USA. June
Caspian Tern (Sterna caspia) in flight, diving for prey. Lake Ellesmere, Canterbury, South Island, New Zealand
Two Red-footed booby (Sula sula) perched on branches, Genovesa Island, Galapagos Islands, Ecuador
White tailed eagle (Haliaeetus albicilla) calling in flight, Isle of Mull, Argyll Scotland, UK. October
Common buzzard (Buteo buteo) in flight with hoar frost. Northamptonshire, UK, January
Golden eagle (Aquila chrysaetus), Kuusamo, Finland, January
Common loon (Gavia immer) nesting at edge of lake and calling, Acadia National Park, Maine, USA, August
RF- Carrion crow (Corvus corone) flying from a snow covered pine tree in a winter landscape. Valsavarenche, Gran Paradiso NP, Italy, March
Clarks Grebe (Aechmophorus clarkii) with yawning chick riding on back, Bear River Refuge, Utah, July
Ptarmigan (Lagopus muta) male on snowy ridge on Cairngorm mountain, Cairngorms National Park, Scotland, UK, March
White-throated dipper (Cinclus cinclus) winter with river flooding, Kuusa, Laukaa, Keski-Suomi, Lansi- ja Sisa-Suomi / Central and Western Finland, Finland. January
Pheasant (Phasianus colchicus), plumage detail. UK, March
Violet sabrewing hummingbird (Campylopterus hemileucurus) male hummingbirds territorial fight at dawn, Turrialba, Costa Rica
Great crested grebe (Podiceps cristatus) pair performing courtship displaying at dawn, backlit and surrounded by mist, Cheshire, UK, April
Northern gannets (Morus bassanus) in flight in Force 8 gales above raging seas. Shetland Islands, Scotland, UK, September
Gentoo penguins (Pygoscelis papua) group walking along snow, Cuverville Island. Antarctic Peninsula, Antarctica
Raven (Corvus corax) calling in the snow, Kemijarvi, Finland, February
Red-and-Green Macaws (Ara chloropterus), feeding at the wall of a riverside clay lick. Blanquillo Clay Lick, Manu Biosphere Reserve, Peru. November
Mute swan (Cygnus olor) parents and cygnets on a misty lake at sunrise. London, UK. May
Sword-billed hummingbird (Ensifera ensifera) hovering in flight, North-Ecuador
Great spotted woodpecker (Dendrocopos major), Scotland, November
RF- Adelie Penguin (Pygoscelis adeliae) on iceberg. Yalour Islands, Antarctic Peninsula, Antarctica. February
Lady Amhersts pheasant (Chrysolophus amherstiae) Kanding, Sichuan, China
Tawny owl (Strix aluco) young sitting in tree amongst autumn leaves, Czech Republic, November. Captive
Shoebill stork (Balaeniceps rex) female feeding on a Spotted African lungfish (Protopterus dolloi) in the swamps of Mabamba, Lake Victoria, Uganda.. Sequence 2/13
Starling (Sturnus vulgaris) feeding fledgling chicks in urban garden. Greater Manchester, UK. May
Resplendent quetzal (Pharomachrus mocinno), captive, Chiapas, southern Mexico, IUCN Near Threatened, April
Shoebill stork (Balaeniceps rex) feeding on a Spotted African lungfish (Protopterus dolloi) in the swamps of Mabamba, lake Victoria, Uganda
Umbrella cockatoo (Cacatua alba), portrait. Captive