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Tiger (Panthera tigris) camouflaged amongst tall grass, looking back at photographer. Assam, India, January
Large aerial pitchers of Veitchs pitcher plant (Nepenthes veitchii) growing up a tree trunk. Maliau Basin, Borneo
Emma Grays Forest Lizard (Calotes emma) in threat display, Xishuangbanna National Nature Reserve, Yunnan Province, China, March
Bornean horned frog (Megophrys nasuta) amongst leaf-litter on forest floor. Danum Valley, Sabah, Borneo, Malaysia
Giant panda (Ailuropoda melanoleuca) in snow, captive
Melanistic leopard / Black panther (Panthera pardus) on territorial patrol on track, Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India
Detail of rhinopores and the colour pattern of a toxic Nudibranch (Goniobranchus coi) on a coral reef. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Spot-bellied eagle owl (Bubo nipalensis) perched on branch, Jim Corbett National Park, Uttarakhand, India
Mandarinfish (Synchiropus splendidus) pair spawning. The larger male (behind) is lifting the female into the spawning rise on his pectoral fin, the smaller female (foreground) is releasing eggs
Rusty-spotted cat (Prionailurus rubininosus phillipsi) captive, occurs in Sri Lanka
Bamboo forest, Shikoku Mura open air museum, Takamatsu, Shikoku Island, Japan, May
A large herd of Tien Shan Argali (Ovis ammon karelini) making tracks through snow. Naryn National Park, Kyrgyzstan, Central Asia, November
Wolverine (Gulo gulo) climbing birch tree, Kamchatka, Far East Russia, April
Small herd of Asiatic Wild Ass (Equus hemionus) running, Gobi National Park, Mongolia
Male Carolas Parotia (Parotia carolae) bird of paradise displaying on his display court to five females perched above, Crater Mountain Wildlife Management Area, Eastern Highlands Province
Pigtail Macaques (Macaca nemestrina) crossing purpose-built primate wires"Pigtail Macaques (Macaca nemestrina) crossing purpose-built " primate wires" connecting isolated patches of forest, Kinabatangan River, Sukau, Sabah, Borneo, September 2008
Sumatran orangutan (Pongo abelii) female with very young baby, Gunung Leuser National Park, Sumatra, Indonesia
Long-tailed macaque (Macaca fascicularis) in a fruiting strangler fig tree (Ficus dubia). Gunung Palung National Park, Borneo
Tualang / Mengaris tree (Koompassia excelsa) emerging from canopy amongst cloud. Danum Valley, Sabah, Borneo, May 2011
Cup fungi (Cookeina sp) growing on decaying log, Danum Valley, Sabah, Borneo
Pitcher plant (Nepenthes veitchii x stenophylla), a natural hybrid. Maliau Basin, Borneo
Komodo Dragon (Varanus komodoensis) in habitat, Komodo National Park, Indonesia
Aerial view of the River Kinabatangan and riverine tropical rainforest, Sabah, Malaysia, Borneo, Asia
Mist and low cloud hanging over lowland rainforest, just after sunrise, with Menggaris Tree (Koompassia excelsa) prominent in the foreground. Danum Valley, Sabah, Borneo
Great egret (Ardea alba) flying across lake, Ranthambhore National Park, India
Terskey Alatau Mountains, Karakol, Kyrgyzstan. August 2016
Young Japanese macaque (Macaca fuscata) kissing another macque in the thermal hot springs, Jigokudani, Japan, January
Orang utan (Pongo pygmaeus pygmaeus) mother with baby climbing on her back, occurs in Borneo, captive, Endangered
Japanese macaque / Snow monkey {Macaca fuscata} mother grooming four-day-old newborn baby, Jigokudani, Nagano, Japan
Four subadult giant pandas (Ailuropoda melanoleuca) climbing in a tree, Wolong Nature Reserve, China, captive
Loggerhead sea turtle (Caretta caretta) emerging from shell, Dalyan delta, Turkey, July
Indian fox (Vulpes bengalensis) pups at play by a den in the Little Rann, Kutch, Gujarat, India. Honorable mention in the Land Mammals category of the BigPicture Competition 2014
Japanese Macaque (Macaca fuscata) baby enjoying a relaxing moment in the hot spring in Jigokudani, Japan. January
Thimble jellyfish (Linuche unguiculata) aggregation, My Paradise Reef, Cabilao Island, Bohol, Central Visayas, Philippines, Pacific Ocean
Three False clownfish (Amphiprion ocellaris) in Sea anemone (Heteractis magnifica) Lighthouse Reef, Cabilao Island, Bohol, Central Visayas, Philippines, Pacific Ocean
RF- Sumatran tiger (Panthera tigris sumatrae) with cub, aged four months, captive, occurs in Sumatra, Indonesia
Juvenile Sumatran tiger (Panthera tigris sumatrae), aged four months, captive, occurs in Sumatra, Indonesia
Bengal Tiger (Panthera tigris tigris) family, Pench National Park, Madhya Pradesh, India, taken on location for Tiger - Spy in the Jungle December 2006
Bengal tiger (Panthera tigris tigris) portrait of a 19-month male. Bandhavgarh National Park, Madhya Pradesh, India
Bengal tiger portrait {Panthera tigris tigris} Bandhavgarh NP, Madhya Pradesh, India
Tiger {Panthera tigris} head profile, Bandhavgarh NP, India
RF- Young male Bengal tiger, head portrait in profile (Panthera tigris tigris). Bandhavgarh National Park, India
Whooper swans (Cygnus cygnus) group of three, camouflaged on frozen lake, Kussharo, Hokkaido Japan
Japanese / Red-crowned crane (Grus japonicus) three in flight, Hokkaido Japan February
Palm (Areaceae) leaves silhouetted in mist at dawn, Royal Chitwan National Park, Nepal
White throated kingfisher (Halcyon smyrnensis) perched on a reed, Sunderban tiger reserve, West Bengal, India
Drifting sand dunes in the Gobi desert, near Dunhuang, Gansu, China
Drifting sand dunes in the Gobi desert, near Dunhuang in Gansu, China