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Sperm whales (Physeter macrocephalus) large group gathering together, rubbing together to shed skin, interacting and defecating. Indian Ocean, March
Hanuman langur (Presbytis entellus) juvenile, running up tree trunk, Bandhavgarh National Park, India
Black-naped monarch / Black-naped blue flycatcher (Hypothymis azurea) male, with insect prey in beak, feeding three chicks at nest, Wilpattu National Park, Sri Lanka
Three Sri Lankan elephants (Elephas maximus maximus) with two calves walking along forest road, Yala National Park, Southern and Uva Provinces, Sri Lanka
Two Millipedes (Julidae) close up portrait, Mulu, Sarawak, Malaysia
Great anglehead lizard (Gonocephalus grandis) resting on tree trunk, Mulu, Sarawak, Malaysia
Black-headed cat snake (Boiga nigriceps) coiled, portrait, Kubah National Park, Sarawak, Malaysia
Sri Lankan leopard (Panthera pardus kotiya) resting on tree branch, Yala National Park, Southern and Uva Provinces, Sri Lanka
Two Chestnut-headed bee-eaters (Merops leschenaulti) perched on branch, Yala National Park, Southern and Uva Provinces, Sri Lanka
Common ghostgoby (Pleurosicya mossambica) camouflaged on a hard coral colony (Plesiastrea sp)Common ghostgoby (Pleurosicya mossambica) camouflaged on a hard coral colony (Plesiastrea sp.), Ambon Bay, Maluku Archipelago, Indonesia. Banda Sea
Two Pallas's cat (Otocolobus manul) kittens waiting at their den for their mother to return from hunting, Mongolia, June, 2017. Filmed for BBC series Big Cats
A female Pallas's cat (Otocolobus manul) with two young kittens hiding amongst rocks at their den site, Mongolia, June, 2017. Filmed for BBC series Big Cats
Pair of Coleman shrimps (Periclimenes colemani) living in a fire urchin (Asthenosoma varium). The female is the larger shrimp
Pair of Broadclub cuttlefish (Sepia latimanus) mating in a coral reef, Kagoshima Prefecture, Japan. Having succeeded in gaining the female's acceptance
Male Siberian flying squirrel (Pteromys volans orii) attempting to enter nest with female inside during reproductive season but stopped mid-way then rejected. Hokkaido, Japan. March
Composite image showing Siberian flying squirrel (Pteromys volans orii) defecating for duration of about twenty seconds after emerging from nest to forage. Hokkaido, Japan. March
Split-second prior to Siberian flying squirrel (Pteromys volans orii) landing on tree, adjusting its attitude in mid-flight toward relatively vertical position to reduce speed
Leaves of Momiji Japanese maple trees (Acer) turning red in autumn. Kyoto, Japan
Autumn foliage of Momiji Japanese maple trees (Acer). Kyoto, Japan
Pair of Siberian flying squirrel (Pteromys volans orii) sharing nest in tree hole in preparation for reproductive season. Hokkaido, Japan. March
Spawning Leaf corals (Pavona decussata). Spawning is synchronized over a large area with many coral colonies, Kyushu, Japan, Pacific Ocean. July
Reef manta ray (Mobula alfredi) swimming with a Twinspot snapper (Lutjanus bohar), Okinawa, Japan, Pacific Ocean
Female Northern pygmy squid (Idiosepius paradoxus) preparing to lay another egg on a blade of eelgrass (Zostera marina), Yamaguchi Prefecture, Japan
Pair of Japanese dwarf flying squirrels (Pteromys volans orii) during the reproductive season, on branch nlooking down, Hokkaido, Japan. The female is on the left, male right
Long-tailed tit (Aegithalos caudatus) feeding on icicle formed from sap of Painted maple tree (Acer pictum). Hokkaido, Japan. February
Male Korean seahorse (Hippocampus haema) giving birth. Kumamoto Prefecture, Kyushu, Japan
Spinner dolphin (Stenella longirostris) juvenile surfacing next to its mother, Sri Lanka
Tufted gray langur (Semnopithecus priam thersites subspecies) sitting in nook f tree, Trincomalee, Sri Lanka, Endangered on the IUCN Red List
Thorny seahorse (Hippocampus histrix) making its way across the muck and rubble substrate of Lembeh Strait in North Sulawesi, Indonesia
Sailfin tang (Zebrasoma veliferum) on coral reef at night, Indonesia, Indo-Pacific
Nudibranch (Goniobranchus geminus) on a coral reef, dorsal view, portrait, Similan Islands, Thailand, Andaman Sea
Longhorn cowfish (Lactoria cornuta) juvenile, swimming over seabed at night, Bitung, North Sulawesi, Indonesia, Molucca Sea
Common clownfish (Amphiprion ocellaris) in the tentacles of its host, Magnificent sea anemone (Heteractis magnifica), on a coral reef, Bitung, North Sulawesi, Indonesia, Molucca Sea
Nudibranch (Chromodoris annae) on a coral reef, Bitung, North Sulawesi, Indonesia, Molucca Sea
Pair of Nudibranchs (Flabellina exoptata) just prior to mating, the mating apparatus of the one on left is visible, Bitung, North Sulawesi, Indonesia, Molucca Sea
Nudibranch (Phyllidia varicosa) portrait on white background, Bitung, North Sulawesi, Indonesia
Large school of Glassfish (Apogonidae) swimming over colourful corals (Dendronephthya sp)Large school of Glassfish (Apogonidae) swimming over colourful corals (Dendronephthya sp.) on a reef, Similan Islands, Thailand, Andaman Sea
Warty frogfish (Antennarius maculatus) yawning in its hiding place amongst white sponges, Bitung, North Sulawesi, Indonesia, Molucca Sea
Leopard (Panthera pardus) hiding in dense foliage, Bardia National Park, Terai, Nepal
Bengal tiger (Panthera tigris tigris) walking through river, snarling, Bardia National Park, Terai, Nepal. Endangered
Green bee-eater (Merops orientalis) calling, portrait, Bardia National Park, Terai, Nepal
Greater one-horned rhinoceros, (Rhinoceros unicornis) walking through shallow water, Bardia National Park, Terai, Nepal
Changeable hawk-eagle / Crested hawk-eagle (Nisaetus cirrhatus) perched on branch, Bardia National Park, Terai, Nepal
Spotted deer (Axis axis), small herd leaping through grass, Bardia National Park, Terai, Nepal
Asian elephant (Elephas maximus indicus) male, eye detail through foliage, Bardia National Park, Terai, Nepal
Greater one-horned rhinoceros (Rhinoceros unicornis) standing in shallow water watching a White-throated kingfisher (Halcyon smyrnensis) fly by, Bardia National Park, Terai, Nepal
Grey langurs (Semnopithecus entellus) family group, sitting and climbing in tree, Bardia National Park, Terai, Nepal
Spotted deer (Axis axis) herd in the forest, with just their antlers visible. Bardia National Park, Terai, Nepal