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North American Bison (Bison bison) coated in frost standing on snow, Yellowstone National Park, Wyoming, USA, January
Gemsbok (Oryx gazella) in sand dunes, Namibia. Finalist in the Mammals Category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
RF - Bison (Bison bison) breathing in the cold air, Yellowstone National Park, USA, February
Llama {Lama glama} portrait, captive, Peru
American bison (Bison bison) female covered in hoar frost near hot spring, portrait. Midway Geyser Basin, Yellowstone National Park, USA. February
Highland cow lying on Heather, Curbar Edge, Peak District National Park, Derbyshire. August
Herd of American bison (Bison bison) in snow, Yellowstone National Park, Wyoming, Yellowstone, January
Fallow deer (Dama dama) grazing, Holkham, Norfolk, November 2011
Red deer (Cervus elaphus) stag in the snow, Scotland, March
Reindeer (Rangifer tarandus) herd, antlers in velvet, walking across upland moor, Cairngorms, Scotland, UK, August
Masai giraffe {Giraffa camelopardalis} mother nuzzling baby, Lower Masai Mara GR, Kenya
Red deer (Cervus elaphus) stag in upland landscape. Lochcarron, Highlands, Scotland, UK
Male Saiga antelope (Saiga tatarica) in winter, The Black Lands National Park, Kalmykia, Russia
Okapi (Okapia johnstoni). Beauval Zoo Parc, France. Captive
Red deer (Cervus elaphus) backlit at dawn with visible breath. UK. October
Bearded pig (Sus barbatus) digging in sand, foraging for crabs on beach, Bako National Park, Sarawak, Borneo
Bearded pig (Sus barbatus) close up of snout, Bako National Park, Sarawak, Borneo
Alpaca portrait {Lama pacos} Altiplano, Bolivia
Caribou / Reindeer bull (Rangifer tarandus) close-up grazing, Denali National Park, Alaska, USA, September
Moose (Alces alces) bull swimming in water, Baxter State Park, Maine, USA
American bison (Bison bison) Yellowstone National Park, Montana, USA
Bison (Bison bison) in snowstorm. Yellowstone National Park, USA, February
Roe deer fawn resting portrait {3-weeks-old} {Capreolus capreolus} Grampian, Scotland, UK
Alpine ibex (Capra ibex) female, Jura, Switzerland, September
Guanaco (Lama guanicoe), Punta Norte, Peninsula Valdes, Argentina
Llama (Lama glama), Los Glaciares National Park, Argentina
RF - Red Deer stag (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Muskox (Ovibos moschatus) with snow on face, Dovrefjell National Park, Norway, February 2009. WWE INDOOR EXHIBITION
Red deer (Cervus elaphus) in Richmond Park with Roehampton Flats in background, London, England, UK, September. 2020VISION Book Plate
Wild boar (Sus scrofa) female, Forest of Dean, Gloucestershire, UK, March
Red deer (Cervus elaphus) stag silhouetted at sunset, Scotland, UK, February
Bison (Bison bison) herd walking in snow, Yellowstone National Park, Wyoming, USA, February
Red deer stag (Cervus elaphus) in heavy snow, Cheshire, UK, March
Siberian musk deer (Moschus moschiferus) male in snow, Irkutsk, Russia. January
Impala with Oxpeckers. Kruger National Park, South Africa
American bison (Bison bison) male walking through deep snow. Hayden Valley. Yellowstone National Park, Wyoming, USA. January
Alpine ibex (Capra ibex), male in habitat, Bernese Alps, Switzerland. July
European bison (Bison bonasus) behind tree, Bialowieza NP, Poland, February
Bison (Bison bison) in snow. Yellowstone National Park, USA, February
Dromedary camel (Camelus dromedarius) train crossing Merzouga dunes, Tafilalt, Sahara, Morocco, March 2007
Highland cattle bull portrait {Bos taurus} Scotland UK
Bison (Bison bison) female with calf walking through snow in front of frost-covered forest, Yellowstone National Park, USA
Gaur (Bos gaurus) Manas National Park UNESCO World Heritage Site, Assam, India
Red Deer stag and hind (Cervus elaphus) silhouetted at sunset, Holkham Park, Norfolk, UK, December
Aerial view of Reindeer (Rangifer tarandus) fording water whilst being herded by Nenet herders. Nenets Autonomous Okrug, Arctic, Russia, July 2017
Red Deer (Cervus elaphus) stag seen from behind a tree. Cairngorms National Park, Scotland, February
Roe Deer stag (Capreolus capreolus) standing in a woodland glade. Potterick Carr Nature Reserve, Doncaster, Yorkshire, England, UK, July
Two Red deer (Cervus elaphus) stags, fighting during the rut, Cheshire, UK. July
Red deer stag (Cervus elaphus) during rut on lookout, Exmoor National Park, Somerset / Devon, England. October
Female American buffalo (Bison bison) walking across grassland with newborn calf, Yellowstone National Park, Wyoming, USA. May
Male Nubian Ibex (Capra nubiana) at the Arabian Wildlife Centre captive-breeding project, Sharjah, United Arab Emirates. (captive)
Male Masaai Giraffe {Giraffa camelopardalis} licking nose, Serengeti NP, Tanzania
Elk (Cervus canadensis) bull and cw sniffing noses, Yellowstone National Park, Wyoming, USA. September
Rocky mountain elk (Cervus elaphus nelsoni) bull bugling during rut. Rocky Mountain National Park, Colorado, USA. October
Alpine ibex (Capra ibex) resting in mountain pasture, Ticino, Switzerland
Tibetan gazelle / Goa (Procapra picticaudata) Keke Xili / Hoh Xil Nature Reserve, Tibetan High plateau, Qinghai, China
Guanaco (Lama guanicoe) in grassland with Torres del Paine rock towers in background. Torres del Paine National Park, Patagonia, Chile. January 2020
Hippopotamus (Hippopotamus amphibius) with water hyacinth still on back after leaving water, Mana Pools National Park, Zimbabwe
Autumn quaking aspen grove (Populus tremuloides) with trunks scarred by browsing elk (Cervus canadensis). Below Turret Ridge, Uncompahgre National Forest, Colorado, USA. October
White-tailed deer (Odocoileus virginianus) doe standing in snow, New York, USA
Altai argali sheep (Ovis ammon), three grazing on slope. Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China. October
Tibetan gazelle (Procapra picticaudata) in snow. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October
Chital (Axis axis) female leaping. Tadoba Andhari Tiger Reserve / Tadoba National Park, Maharashtra, India
Mongol driving a camel caravan of Bactrian camel (Camelus bactrianus) Bashang Grassland, Zhangjiakou, Hebei Province, Mongolia, China
Red deer (Cervus elaphus) silhoutted at dusk, Richmond Park, London, England, UK. October
Moose bull (Alces alces) walking in forest clearing, Denali National Park, Alaska, USA, September
Moose (Alces alces) bull swimming across river. Grand Teton National Park, Wyoming, USA. September
Guanacos (Lama guanicoe), two males fighting, trying to over power opponent and bite each others testicles. Torres del Paine National Park, Patagonia, Chile. December
Silver Dik-dik (Madoqua piacentinii) portrait, Masai Mara, Kenya. March
Guanaco (Lama guanicoe) female, alert whilst looking at Puma (Puma concolor) on on opposite slope, portrait. Torres del Paine National Park, Patagonia, Chile. November
Gayal / Mithun (Bos frontalis) on river bank. Arunachal Pradesh, North East India. November 2014
Chinese Water Deer (Hydropotes inermis) Norfolk UK September
Roe deer (Capreolus capreolus) buck with vegetation on antlers in rutting season, Scotland, UK. August
Alpine ibex (Capra ibex) adult males fighting in alpine landscape, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Alpine ibex (Capra ibex) struggling in deep snow on a steep slope, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, January
Snow leopard (Uncia uncia) with Bharal (Pseudois nayaur) prey, Serxu County, Garze Prefecture, Sichuan Province, China
Pere Davids deer / Milu (Elaphurus davidianus) during the rutting season, stag, Hubei Tian ezhou Milu National Nature Reserve, Shishou, Hubei, China
Vicuna (Vicugna vicugna), five running across salt flat with mountain in background. Salar de Ascotan, Chile. September 2018
Red deer (Cervus elaphus) stag, Saxony, Germany. September
Bactrian camels (Camelus bactrianus) on sand, Nubra Valley, Ladakh, India. September
Muskox (Ovibos moschatus), male portrait. Dovrefjell National Park, Norway
Male American Bison (Bison bison) walking through snow. Firehole River Valley, Yellowstone National Park, Wyoming, USA. January
Sable antelope (Hippotragus niger) bull charging through the shallows of the Selinda Spillway, with another behind, northern Botswana, August
