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Two Asian weaver ants (Oecophylla smaragdina), queens founding a new nest, tending to eggs, West Bengal, India
Namib desert dune ant (Camponotus detritus), queen on sand with visible wing scars on her thorax, Swakopmund, Namibia
Cardinal jumper (Phidippus cardinalis) spider on a leaf, Florida, USA. Controlled conditions
Yellow-tail moth (Euproctis similis) on a leaf, Montiaghs Moss National Nature Reserve, County Antrim, Northern Ireland. July
Digital composite of Swallowtail butterfly (Papilio machaon) caterpillar, pupa, chrysalis and emerging adult
Glasswing butterfly (Greta oto) resting on leaf. Captive
RF - Common darter dragonfly (Sympetrum striolatum) resting on Foxglove (Digitalis purpurea) with dew droplets on wings. Cornwall, England, UK. June
Botany Bay diamond weevil (Chrysolopus spectabilis) close up, Toowoomba, Queensland, Australia
Firefly (Lamprohiza splendidula) light trails of males in forest at dusk, Bavaria, Germany. July
Mexican Fiddler Crab (Leptuca crenulata) male displaying, Morua Estuary, near Rocky Point, Gulf of California, Mexico, June
Hermit crab (Dardanus calidus) Vis Island, Croatia, Adriatic Sea, Mediterranean
Cairns birdwing butterfly (Ornithoptera euphorion) male resting on fern, large butterfly endemic to wet tropics in far north Queensland, Cairns, Queensland, Australia
Lamington spiny crayfish (Euastacus sulcatus) blue form, in leaf litter under tree roots with claws raised, rare crayfish that commonly forages away from water on the forest floor above 300m
Galapagos sea lion (Zalophus wollebaeki) inquisitive pup playing with Sally Lightfoot Crab (Grapsus grapsus). Plazas Island, Galapagos, Ecuador
African golden emperor moth (Gonimbrasia krucki), close up of male head, Kenya, Africa Controlled conditions
White-ringed atlas moth (Epiphora mythimnia) portrait, Kenya, Africa
Taiwanese squeaking silkmoth (Rhodinia verecunda) controlled conditions
Giant house spider (Eratigena atrica) close up showing large mandibles, Banbridge, County Down, Northern Ireland. September
Thorny devil stick insect (Eurycantha calcarata), Willaumez Peninsula, New Britain, Papua New Guinea, December
Spider wasp (Pompilidae) with spider prey, Buxa Tiger Reserve, India
Sallow kitten moth (Furcula furcula), Peak District National Park, Derbyshire, UK. August. Focus stacke image
Longhorn orb web spider (Macracantha arcuata), Danum Valley, Sabah, Borneo
Lantern bug (Pyrops whiteheadi), Danum Valley, Sabah, Borneo
Swimming crab (Cronius ruber) Canary Islands, Tenerife. Invasive species
Termite hill and acacia tree, Shaba National Reserve, North Kenya
Rosemary beetle (Chrysolina americana) feeding on Lavender (Lavandula angustifolia) in garden Cheshire, UK, June
Bordered patch butterfly (Chlosyne lacinia) on Indian blanket (Gaillardia pulchella). Hill Country, Texas, USA
Four-spotted chaser dragonfly (Libellula quadrimaculata), two resting on reeds. Cornwall, England, UK. May
Apollo butterfly (Parnassius apollo) female laying eggs on branch. Parainen, Finland. July
Wolf spider (Pardosa amentata) close up, Leeds, Yorkshire, England, UK, April. Focus stacked image
RF - Small tortoiseshell butterfly (Aglais urticae) nectaring on Verbena (Verbena sp). Cornwall, England, UK. September. (This image may be licensed either as rights managed or royalty free.)
Atlantic blue crab (Callinectes sapidus) hunting in Turtlegrass (Thalassia testudinum) seagrass bed, view towards sky. Florida Keys, Florida, USA
Lord Howe Island stick insect (Dryococelus australis), Bristol Zoo Gardens, Bristol, UK
Apple hawk-moth (Langia zenzeroides formosana) Nzaus Mamei Jianshih Village, Hsssinchu County, Taiwan, Controlled conditions
Crab spider (Thomisus onustus) on white flowerhead, Sierra de Grazalema Natural Park, southern Spain, May
Spoonwing / Thread-winged antlion (Nemoptera bipennis) on flower, Sierra de Grazalema Natural Park, southern Spain. June
Cockchafer beetle (Melolontha melolontha) on lichen encrusted twig. France. May
Firefly (Lamprohiza splendidula) light trails in woodland at night, multiple exposure. The Netherlands. June
Portrait of a Emerald damselfly (Lestes sponsa) Yorkshire, August. Focus stacked image
Stag beetle (Lucanus cervus), adult male on oak tree, Italy
Black-veined white (Aporia crataegi) butterfly pair, in flight and nectaring on Thistle. Jyvaskyla, Central Finland. June
RF - Male Comet Moth (Argema mittrei). Mantadia National Park, eastern Madagascar. (This image may be licensed either as rights managed or royalty free.)
Brimstone butterfly (Gonepteryx rhamni) resting among foliage, Meeth Quarry, Devon, UK. June
Marbled White butterfly (Melanargia galathea) resting on flower head, Dunsdon Nature Reserve, Devon, UK. July
Male common blue butterfly (Polyommatus icarus) basking wings open on grass, Vealand Farm, Devon, UK. June
Glasswing butterfly (Greta oto), Mariposario del Drago, Icod de Los Vinos, Tenerife, August
Painted lady butterfly (Vanessa cardui) feeding on knapweed Sutcliffe Park Nature Reserve, Eltham, London, England, UK. June
Inser Weiher in autumn, a pond with red coloured water, caused by dissolved tannins. Berner Seeland, Berne, Switzerland, November. Photographed for The Freshwater Project