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Red fox (Vulpes vulpes) adult hunting for rodents by snow diving in deep snow. Hayden Valley, Yellowstone, USA. January
Boky-boky (Mungotictis decemlineata) or Narrow-striped mongoose adult foraging in leaf litter. Dry deciduous forest, Kirindy, western Madagascar
Fosa or Fossa (Cryptoprocta ferox)(sometimes erroneously fossa) male in dry deciduous forest. Kirindy, western Madagascar
Puma (Puma concolor puma), female sitting in grassland. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. November
Guanacos (Lama guanicoe), two males fighting, trying to over power opponent and bite each others testicles. Torres del Paine National Park, Patagonia, Chile. December
Puma (Puma concolor puma), young male crossing road in front of car. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December 2018
Bush cricket / Katydid (Tettigoniidae ), female leaf mimic ovipositing into branch. Manu Biosphere Reserve, Peru
Eyelash pit viper (Bothriechis schlegelii) (Viperidae: Crotalinae). Distictive yellow / orange oropel form. Arboreal species resting on wild ginger flower in mid-altitude rainforest under storey
Proboscis monkey (Nasalis larvatus) Riverine forest, Kinabatangan River, Sukau, Sabah, Borneo
Adult Western / Horsfields tarsier (Tarsius bancanus) in forest understorey at night. Lowland dipterocarp rainforest, Danum Valley, Sabah, Borneo
RF- Wild male Jaguar (Panthera onca palustris) running through shallows of backwater of Cuiaba River in late afternoon sunlight. Northern Pantanal, Brazil
Giant River Otter (Pteronura brasiliensis) feeding on Striped Catfish or Cachara (Pseudoplatystoma fasciatum) amongst water hyacinth. Tributary off the Cuiaba River, Mato Grosso, Pantanal, Brazil
Siberian flying squirrel (Pteromys volans orii) feeding on Sakhalin fir tree. Hokkaido, Japan. March
North Pacific humpback whale (Megaptera novaeangliae kuzira) breaching at sunset, Kupreanof Island, Frederick Sound, Alaska, USA, July
Blue sea slug (Glaucus atlanticus) that was washed ashore with a mass, multi-day stranding of thousands of Indo-Pacific Portuguese man-of-war (Physalia utriculus)
Asian sheepshead wrasse (Semicossyphus reticulatus) mature male in breeding colouration. Niigata, Japan
Fireflies (Luciola parvula himebotaru) flashing at night for courtship and reproduction. Gifu, Japan. Composite image
Killer whale / Orca (Orcinus orca) mature male, swimming underwater. Hamn, Senja, Norway, Atlantic Ocean
Humpback whale calf (Megaptera novaeangliae) male resting beneath his resting mother. Vava u, Tonga, South Pacific
Atlantic walrus (Odobenus rosmarus rosmarus) hauled out on ice floe, Svalbard, Norway June
Humpback whales (Megaptera novaeangliae) engaged in courtship, with the male hovering above the female. Vava'u, Tonga, Pacific Ocean
Sperm whale (Physeter macrocephalus) female at the ocean surface with her mouth open. This whale has just surfaced from foraging in deep water. Ogasawara or Bonin Islands, Japan
Twinspot snapper (Lutjanus bohar) shoal rushing up to spawn, releasing a cloud of milky white sperm and eggs in the water column. Caroline Islands, Palau, November
Giant panda (Ailuropoda melanoleuca) cub in tree, Chengdu Panda Breeding Centre, Sichuan, China. Captive
Smalltooth sand tiger shark (Odontaspis ferox), view from below. El Hierro. Canary Islands
Fangtooth moray (Enchelycore anatina) with open mouth, portrait. Tenerife, Canary Islands
Arrow crab (Sternorhynchus lanceolatus). Tenerife, Canary Islands
Short-finned pilot whale (Globicephala macrorhynchus) calf with fetal lines characteristic of newborn specimens, with pod. Tenerife, Canary Islands
Ocean sunfish (Mola mola) and Pilot fish (Naucrates sp). Tenerife, Canary Islands
Napoleon wrasse (Cheilinus undulatus) on a coral reef. Ras Mohammed National Park, Sinai, Egypt. Red Sea
Bluestreak cleaner wrasse (Labroides dimidiatus) cleans among the sharp teeth of a predatory roving coral grouper (Plectropomus pessuliferus) beneath an overhang on a coral reef
Moon jelly (Aurelia aurita) drifts over a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Arctic fox (Alopex lagopus) camouflaged in winter pelage. Svalbard, Norway, April
Polar bear (Ursus maritimus) standing on ice. Svalbard, Norway, July
Roe Deer stag (Capreolus capreolus) standing in a woodland glade. Potterick Carr Nature Reserve, Doncaster, Yorkshire, England, UK, July
RF - Nile crocodile (Crocodylus niloticus head close up with jaws open, Chobe river, Botswana)RF - Nile crocodile (Crocodylus niloticus head close up with jaws open, Chobe river, Botswana. (This image may be licensed either as rights managed or royalty free.)
Lion (Panthera leo) drinking at night, Zimanga private game reserve, KwaZulu-Natal, South Africa
RF - African elephants (Loxodonta africana) in water, trunks touching, Zimanga game reserve, South Africa. (This image may be licensed either as rights managed or royalty free.)
Silver Dik-dik (Madoqua piacentinii) portrait, Masai Mara, Kenya. March
Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May
Reef manta ray (Manta alfredi) shoal filter feeding on plankton concentrated by monsoon currents in Raa Atoll, Vandhoo Thila, Maldives. October
Red fox (Vulpes vulpes) in mid air, snow diving / pouncing whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February
Guanaco (Lama guanicoe) female, alert whilst looking at Puma (Puma concolor) on on opposite slope, portrait. Torres del Paine National Park, Patagonia, Chile. November
Sumatran pit viper (Trimeresurus sumatranus) juvenile on tree trunk, in tropical rainforest. Danum Valley, Sabah, Borneo, Malaysia
Flying fish (Exocoetidae) in Sargasso Sea, Atlantic Ocean
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) swimming over Seagrass (Halophila stipulacea) meadow. Marsa Nabaa, Marsa Alam, Egypt
Siberian weasel (Mustela sibirica) standing in snow. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
Giant river otter (Pteronura brasiliensis) resting on fallen tree trunk above Cuiaba River. Pantanal, Mato Grosso, Brazil