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Two Blood pheasant birds (Ithaginis cruentus) males Baima Snow Mountain Nature Reserve, Yunnan, China. October
Pallass cat (Otocolobus manul) resting, Tibetan Plateau, Qinghai, China
Tibetan sand fox (Vulpes ferrilata), Valley near Yushu, Tibetan Plateau, Qinghai, China
Meerkat (Suricata suricatta) standing and keeping watch while another excavates a burrow in soft sand before disappearing underground for the night. Makgadikgadi Pans, Botswana, April
Blue shark (Prionace glauca) with Pilot fish (Naucrates ductor) off Halifax, Nova Scotia, Canada. July
American / Northern lobster (Homarus americanus) partially hidden under seaweed, Nova Scotia, Canada, July
Black-crowned Central American squirrel monkey (Saimiri oerstedii) sitting on branch, Corcovado National Park, Osa Peninsula, Costa Rica, Vulnerable species
American wigeon (Anas / Mareca americana) nest with seven eggs, Anchorage Provincial Park, Grand Manan Island, New Brunswick, Canada, June
Black capped chickadee (Poecile atricapillus) portrait, Anchorage Provincial Park, Grand Manan Island, New Brunswick, Canada, June
Tetrapod tracks preserved in rock at the Joggins Fossil Cliffs UNESCO World Heritage Site, Bay of Fundy, Nova Scotia, Canada. May 2017
North Atlantic Right Whale (Eubalaena glacialis) tail fluke, in the Bay of Fundy, Canada, September
Puma (Puma concolor) resting, Corcovado National Park, Costa Rica, May
Bairds tapir (Tapirus bairdii) Corcovado National Park, Costa Rica, May. Endangered
Bairds tapir (Tapirus bairdii) walking along beach in Corcovado National Park, Costa Rica. Endangered
Ocelot (Leopardus pardalis) walking, camera trap image, Nicoya Peninsula, Costa Rica
Minke whale ( Balaenoptera acutorostrata) breaching, Bay of Fundy, Canada, July
Ocelot (Leopardus pardalis) portrait. camera trap image, Nicoya Peninsula, Costa Rica
Atlantic puffic, (Fratercula arctica), Machias Seal Island, Bay of Fundy, New Brunswick, Canada, August
Atlantic puffic, (Fratercula arctica) portrait. Machias Seal Island, Bay of Fundy, New Brunswick, Canada, August
Leaf mantis (Choeradodis sp. ) Tortuguero National Park, Costa RicaLeaf mantis (Choeradodis sp.) Tortuguero National Park, Costa Rica
Jaguar (Panthera onca) camera trap image, Tortuguero National Park, Costa Rica
Giant owl butterfly, (Caligo memnon) Tortuguero National Park, Costa Rica
Luna moths (Actias luna) New Brunswick, Canada, June
Eyespots on wing of Giant owl butterfly, (Caligo memnon) Tortuguero National Park, Costa Rica, February 2015
Neotropical river otter (Lontra longicaudis) in water, Nicoya Peninsula, Costa Rica, March 2015
Ivy bee (Colletes hederae) new species to the UK in 2001. Male resting on Blackberry, Oxfordshire, England, UK, September
Harvest mouse (Micromys minutus) climbing among wheat, Hertfordshire, England, UK, August, Controlled conditions
Long beaked common dolphin (Delphinus capensis) pod porpoising Baja California, Mexico
Californian Sealion (Zalophus californianus) in kelp (Macrocystis pyrifera) forest, San Benito del Este Island, Baja California Peninsula Pacific Islands Biosphere Reserve, Baja California, Mexico
Purple hairstreak butterfly (Neozephyrus quercus) female with wings open, Hertfordshire, England, UK, July. Focus Stacked Image
Firecrest (Regulus ignicapilla) singing, Wiltshire, England, UK, April
Ants (Formicidae) attacking newly emerged cicada (Cicadacae), Yasuni National Park, Ecuador. Highly commended in the Invertebrates Category of the Wildlife Photographer of the Year Awards (WPOY)
Essex skipper butterfly (Thymelicus lineola) male visiting flower, , Wiltshire, England, UK, July
Sedge Warbler (Acrocephalus schoenobaenus) Greylake RSPB Reserve, Somerset Levels, England, UK, August
Great pied hoverfly (Volucella pellucens) feeding from bramble flower, Wiltshire, England, UK, July. Bumblebee mimic species
Piraputanga (Brycon hilarii) and small Tetra fish (Hyphessobrycon sp. ) Rio Olhio d AguaPiraputanga (Brycon hilarii) and small Tetra fish (Hyphessobrycon sp.) Rio Olhio d Agua, tributary of Rio da Prata, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul
Atlantic salmon (Salmo salar) jumping up waterfall during spawning migration upstream, Ume River, Sweden. July. Photographed for The Freshwater Project
Flightless cormorant (Phalacrocorax harrisi) pair in courtship Puerto Pajas, Isabela Island, Galapagos
Split level view of Brown pelicans (Pelecanus occidentalis) feeding, Tortuga Bay, Santa Cruz Island, Galapagos
Brown pelican (Pelecanus occidentalis) feeding chicks at nest, Puerto Ayora / Academy Bay, Santa Cruz Island, Galapagos
Four-spotted chaser dragonfly (Libellula quadrimaculata) just emerged. Finland, June
Green sea turtles (Chelonia mydas) on corals, Hawaii
Spoonbill (Platalea leucorodia) carrying nest material, The Netherlands
Andean flamingo (Phoenicoparrus andinus) feeding on shore Laguna Hedionda, Altiplano, Bolivia
Jamess flamingo (Phoenicoparrus jamesi) flock on the shore of Laguna Colorada / Reserva Eduardo Avaroa, Altiplano, Bolivia
Sargassum fish (Histrio histrio) at the sea surface with floating sargassum weed. Hawaii
Painted cardinalfish (Archamia fucata) Yap, Micronesia
Caribbean manatee or West Indian manatee (Trichechus manatus) mother with baby, age two days, captive, Beauval Zoo, France
