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Timber Wolves (Canis lupus) resting on a rock by the shoreline. Near Mussel Inlet, Great Bear Rainforest, British Columbia, Canada, October
Pine marten (Martes martes) climbing Scots pine tree, Scotland, UK. May
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Black-headed gull (Chroicocephalus ridibundus) in flight, artistically blurred photograph, Cheshire, UK, February. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Japanese Macaques (Macaca fuscata) in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
Lioness (Panthera leo) carrying her cub, Masai-Mara Game Reserve, Kenya
The River Thames and Houses of Parliament at Westminster at sunset with gull, UK, November 2005
Nicaraguan giant glass frog (Espadarana prosoblepon) La Selva Field Station, Costa Rica
Grey wolf (Canis lupus) adult greeted by cub, captive, USA
Welsh Terrier, bitch with puppies aged 8 weeks in snowy scene
Bison (Bison bison) herd walking in snow, Yellowstone National Park, Wyoming, USA, February
Pine marten (Martes martes) portrait, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Portrait category of the BWPA (British Wildlife Photography Awards) Competition 2016
Two Rooks (Corvus frugilegus) silhouetted as they perch on a tree branch at their roost site at sunset, Gloucestershire, UK, February
Badger (Meles meles) adult, backlit at sunset, scratching its ears, Derbyshire, UK, May. Winner of the Animal Behaviour category of the British Wildlife Photography Awards (BWPA) Competition 2017
RF - Red Deer stag (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Red deer stag (Cervus elaphus) in heavy snow, Cheshire, UK, March
Atlantic Puffin (Fratercula arctica) one landing among resting group, Sule Skerry, Scotland, UK, July
RF- Western clownfish (Amphiprion oceallaris) in Magnificent sea anemone (Heteractis magnifica). Raja Ampat, West Papua, Indonesia
Blue whale (Balaenoptera musculus) underwater, Coronado Islands, Baja California, Mexico, Eastern Pacific Ocean
Piraputanga (Brycon hilarii) shoal, Rio Olhio d agua, tributary of Rio da Prata, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil
Siberian tiger (Panthera tigris altaica) in snow, captive
Impala with Oxpeckers. Kruger National Park, South Africa
Squid (Abraliopsis atlantica) deep sea species from Atlantic Ocean off Cape Verde. Captive
Atlantic Puffins (Fratercula arctica) pair bill rubbing, part of ritual courtship, Skomer Island, Wales, UK, May
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
Black Bear (Ursus americanus) sleeping in a Sitka Spruce tree (Picea sitchensis) alongside Anan Creek, Alaska, July
Giant panda (Ailuropoda melanoleuca) cub yawning. Yuan Meng, first giant panda ever born in France, age 10 months, Captive at Beauval Zoo, Saint Aignan sur Cher, France
Polar bear cub 3 months (Ursus maritimus) playing in the front of the day den in march. Wapusk National Park. Churchill, Manitoba, Canada
Lion (Panthera leo) cubs playing, Masai-Mara Game Reserve, Kenya. Vulnerable species
Three Sperm whales (Physeter macrocephalus) socialising under the surface, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species
RF- African elephant (Loxodonta africana) at sunset. Chobe National Park, Botswana. Endangered species
Mute swan (Cygnus olor) in early morning light removing its own feathers, during moulting, Valkenhorst Nature Reserve, The Netherlands, Europe. August
Goldfinch (Carduelis carduelis) feeding on teasel in falling snow, Leicestershire, UK, February
Red necked pademelon (Thylogale thetis) with joey peering from pouch, in scrubland at edges of rainforest, small macropod found in eastern Australia, Lamington National Park, Queensland, Australia
Southern cassowary (Casuarius casuarius), adult male with chicks keeping a wary eye on the photographer. Near Bloomfield, north Queensland, Australia, September
Eyrean Grasswren (Amytornis goyderi) in typical hopping motion, Andado Station, Northern Territory, Australia, November
Bearded reedling (Panurus biarmicus) female feeding on reed seeds covered in frost, Espoo, Finland, January
Pletail unicornfish (Naso brevirostris) juvenile, its horn will continue to grow as it gets older. Fiji Pacific Ocean
Tabular iceberg floating in Weddell Sea, iceberg broken away from Larson C ice shelf. Adelie penguin (Pygoscelis adeliae) colony dwarfed on sea ice below. Near Danger Islands, Antarctica
White-tailed eagle (Haliaeetus albicilla) taking off with fish prey, Norway, August
Portrait of a Black-capped chickadee, (Poecile atricapillus) with white background, Block island, Rhode Island, USA. Bird caught during scientific research
Eurasian crag martin (Ptyonoprogne rupestris) Kawakarpo Mountain, Meri Snow Mountain National Park, Yunnan Province, China. May
Grey heron (Ardea cinerea) wading at night, reflected in water. Lincolnshire, UK, September
RF - White-legged damselfly (Platycnemis pennipes) peering over Oxeye daisy (Leucanthemum vulgare) flower. The Netherlands. August
Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek against ancient Manchurian fir tree, Land of the Leopard National Park, Far East Russia, November
Fallow deer (Cervus dama) stags at sunset, Holkham Park, North Norfolk, UK, January
