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Common Loons (Gavia immer) one diving underwater, Michigan, USA. June
Moose bull (Alces alces) walking in forest clearing, Denali National Park, Alaska, USA, September
Black oil beetle (Meloe proscarabaeus) male on grass stem in sand dunes, Gower, Wales, UK, February. Focus stacked image
RF - Oceanic whitetip shark (Carcharhinus longimanus) swims in open waters, close to the surface with sun beams. Egypt. Red Sea. (This image may be licensed either as rights managed or royalty free.)
Javan leopard (Panthera pardus melas) roaring, native to the Indonesian island of Java. Captive. Digital composite
Porto Santo Wolfspider (Hogna schmitzi) Porto Santo island, Madeira Archipelago, Portugal. Endemic
Indri (Indri indri) with young, Palmarium Resrve Ankanin ny Nofy, Littoral rainforest, Madagascar
Great Grey Owl (Strix nebulosa) Kuhmo Finland, March
Sparrowhawk (Accipiter nisus) adult female, Norway, October
Golden jackal (Canis aureus) pup sitting on a grassy verge by a path, Danube Delta, Romania, July
African lion (Panthera leo) and lioness after mating, Masai Mara National Reserve, Kenya
Lion (Panthera leo) male walking. Masai Mara National Reserve, Kenya
Lady Amhersts pheasant (Chrysolophus amherstiae) male displaying, Kanding, Sichuan, China. April
European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition
Polar bears (Ursus maritimus) fighting in water. Beaufort Sea, Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Natures Best Photography Competition 2019
Grey-headed flying-fox (Pteropus poliocephalus) spotting and flaring wings to come in to land on a branch. Yarra Bend Park, Kew, Victoria, Australia. ÔÇ¿November 2019
Common Frog (Rana temporaria), and frogspawn in a pond, Coldharbour, Surrey, England. March. Highly commended in the Animal Portrait category of the BWPA Competition 2019
Hard coral (Porites lutea) with Christmas-tree worms (Spirobranchus giganteus), New Caledonia, Pacific Ocean
Bee-eater (Merops apiaster) male offering wasp as nuptial gift, Hungary. June
Common Cuckoo (Cuculus canorus) perched on gate Surrey, England, UK. April
Starling (Sturnus vulgaris) murmuration at sunset, West Pier, Brighton, England, UK
Timber rattlesnake (Crotalus horridus) females and newborn young at maternity site. Pennsylvania, USA. August
Fosa of Fossa (Cryptoprocta ferox) male walking through dry deciduous forest, Kirindy, western Madagascar
Aye-aye (Daubentonia madagascariensis) adult active and foraging in forest canopy at night. Deciduous forests, Daraina, northern Madagascar. Endangered endemic species
Red fox (Vulpes vulpes) adult hunting for rodents by snow diving in deep snow. Hayden Valley, Yellowstone, USA. January
Boky-boky (Mungotictis decemlineata) or Narrow-striped mongoose adult foraging in leaf litter. Dry deciduous forest, Kirindy, western Madagascar
Fosa or Fossa (Cryptoprocta ferox)(sometimes erroneously fossa) male in dry deciduous forest. Kirindy, western Madagascar
Puma (Puma concolor puma), female sitting in grassland. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. November
Guanacos (Lama guanicoe), two males fighting, trying to over power opponent and bite each others testicles. Torres del Paine National Park, Patagonia, Chile. December
Puma (Puma concolor puma), young male crossing road in front of car. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December 2018
Bush cricket / Katydid (Tettigoniidae ), female leaf mimic ovipositing into branch. Manu Biosphere Reserve, Peru
Eyelash pit viper (Bothriechis schlegelii) (Viperidae: Crotalinae). Distictive yellow / orange oropel form. Arboreal species resting on wild ginger flower in mid-altitude rainforest under storey
Proboscis monkey (Nasalis larvatus) Riverine forest, Kinabatangan River, Sukau, Sabah, Borneo
Adult Western / Horsfields tarsier (Tarsius bancanus) in forest understorey at night. Lowland dipterocarp rainforest, Danum Valley, Sabah, Borneo
RF- Wild male Jaguar (Panthera onca palustris) running through shallows of backwater of Cuiaba River in late afternoon sunlight. Northern Pantanal, Brazil
Giant River Otter (Pteronura brasiliensis) feeding on Striped Catfish or Cachara (Pseudoplatystoma fasciatum) amongst water hyacinth. Tributary off the Cuiaba River, Mato Grosso, Pantanal, Brazil
Siberian flying squirrel (Pteromys volans orii) feeding on Sakhalin fir tree. Hokkaido, Japan. March
North Pacific humpback whale (Megaptera novaeangliae kuzira) breaching at sunset, Kupreanof Island, Frederick Sound, Alaska, USA, July
Blue sea slug (Glaucus atlanticus) that was washed ashore with a mass, multi-day stranding of thousands of Indo-Pacific Portuguese man-of-war (Physalia utriculus)
Asian sheepshead wrasse (Semicossyphus reticulatus) mature male in breeding colouration. Niigata, Japan
Fireflies (Luciola parvula himebotaru) flashing at night for courtship and reproduction. Gifu, Japan. Composite image
Killer whale / Orca (Orcinus orca) mature male, swimming underwater. Hamn, Senja, Norway, Atlantic Ocean
Humpback whale calf (Megaptera novaeangliae) male resting beneath his resting mother. Vava u, Tonga, South Pacific
Atlantic walrus (Odobenus rosmarus rosmarus) hauled out on ice floe, Svalbard, Norway June
Humpback whales (Megaptera novaeangliae) engaged in courtship, with the male hovering above the female. Vava'u, Tonga, Pacific Ocean
Sperm whale (Physeter macrocephalus) female at the ocean surface with her mouth open. This whale has just surfaced from foraging in deep water. Ogasawara or Bonin Islands, Japan
Twinspot snapper (Lutjanus bohar) shoal rushing up to spawn, releasing a cloud of milky white sperm and eggs in the water column. Caroline Islands, Palau, November
Giant panda (Ailuropoda melanoleuca) cub in tree, Chengdu Panda Breeding Centre, Sichuan, China. Captive