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Young Tigress (Panthera tigris), 22 month, being frightened by a dead leaf. Banghavgarh National Park, India, April
Portrait of a wild Horse (Equus caballus). The foothills of southwest Reno, Nevada, USA, May
Wild stallion (Equus caballus) lying down in the sagebrush and yawning. The foothills south of Reno, Nevada, USA, May
Brown Hare (Lepus europaeus) portrait gazing alertly. Perthshire, Scotland, June
Domestic dog, Scottish Terrier / Aberdeen Terrier, studio portrait
Group of South American fur seals (Arctocephalus australis) swimming near the rocky shore. Puerto Deseado, Patagonia, Argentina, March 2008
Grey Wolf (Canis lupus) head portrait of male, sleeping in snow, Captive
Zanzibar Red Colobus (Procolobus kirkii) head portrait, Jozani Chwaka Bay NP, Zanzibar, Tanzania
Old male African lion (Panthera leo) portrait, Sabi Sand Game Reserve, South Africa, June
Laysan albatross (Phoebastria immutabilis) head portrait calling mate, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, March
An adult female California sea lion (Zalophus californianus) head portrait, in a rocky underwater cave. La Paz, Baja California Mexico. Sea of Cortez, East Pacific Ocean. October
Polar bear (Ursus maritimus) head close-up portrait of an adult male, with snowflakes on fur, Barter Island, 1002 area of the Arctic National Wildlife Refuge, Alaska
Small Munsterlander head portrait
Dalmatian, head portrait, male aged 4 years
Alaskan Malamute portrait of blue coated adult
Adult male Bornean orangutan (Pongo pygmaeus) in rainforest canopy, Gunung Palung National Park, Borneo, West Kalimantan, Indonesia
Checkered Garter Snake (Thamnophis marcianus marcianus) coiled with tongue protruding. Fennessey Ranch, Refugio, Corpus Christi, Coastal Bend, Texas Coast, USA
Portrait of German Shorthair Pointer in thicket of pink berries, Illinois, USA
Head portrait of black and tan smooth coated Dachshund with zinnias, Illinois, USA
Head portrait of black and white domestic cat (Felix catus) looking up
Close up head portrait of melanistic / black Leopard (Panthera pardus) Captive
Head portrait of male Mandrill (Mandrillus sphinx) Captive. Apenheul zoo; the Netherlands
Red grouse {Lagopus lagopus scoticus} on heather moorland, Derwent Edge, Peak District National Park, UK, September
Eastern green mamba {Dendroaspis angusticeps} strike pose, captive, from East Africa
Grizzly bear (Ursus arctos horribilis) catching salmon in Brooks river, Katmai National Park, Alaska, USA, July
Two Mute swan (Cygnus olor) cygnets, Dorset, UK