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African wild dogs (Lycaon pictus) juveniles playing with the leg of an impala, trying to drag it in three different directions. Mkuze, South Africa
Dugong (Dugong dugon) male feeding on a seagrass meadow (Halophila stipulacea), accompanied by a young Golden trevally (Gnathanodon speciosus). Marsa Nabaa, Marsa Alam, Egypt. Red Sea
Barn owl (Tyto alba) in flight, hunting, Hampshire, England, UK. Captive
Blue-footed booby (Sula nebouxii) courting pair, South coast, Santa Cruz Island, Galapagos
Humpback whale (Megaptera novaeangliae) breaching, near Hout Bay, South Africa
Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October
Killer whale (Orcinus orca) male named Mel attacking young South American sea lion (Otaria flavescens) on beach, Punta Norte, Peninsula Valdes, Argentina
A split level digital composite showing a Basking shark (Ceterhinus maximus) feeding on plankton around St Michaels Mount, Cornwall, UK. June
Common starling (Sturnus vulgaris) murmuration, flock pursued by Peregrine falcon (Falco peregrinus) before landing at winter roost. The Netherlands. February 2020
Male Pileated Woodpecker (Dryocopus pileatus) with beetle larva in beak about to feed two chicks, New York, USA, May
Leaf cutter ants (Atta sp) carrying colourful plant matter, reflected in water, Laguna del Lagarto, Santa Rita, Costa Rica
RF - Lions (Panthera leo) - two brothers patrolling territorial boundary. Short grass plains on the border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Atlantic Puffins (Fratercula arctica) bill rubbing, Skomer Island, Wales, UK, April
Polar bear (Ursus maritimus) young bear rolling around in the snow, on newly formed pack ice during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Red fox (Vulpes vulpes) cubs playing, Vosges, France, May
Pipistrelle bat (Pipistrellus pipistrellus) flying low over water. Surrey, England, August
Caribbean Flamingo (Phoenicopterus ruber) chick under the wing of protective parent, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico
Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April
Male Leach's sea star (Leiaster leachi) broadcast spawning, releasing streams of sperm from its arms. Kagoshima Prefecture, Kyushu, Japan
Great hammerhead shark (Sphyrna mokarran) mouth wide open, feeding in shallow water, surrounded by fish. South Bimini Island, Bahamas
Working English Springer Spaniel puppy, 6 weeks, in playful stance
Mating pair of chalkhill blue butterflies (Lysandra coridon) with wings closed resting on Devils-bit scabious (Succisa pratensis), Hatch Hill, Somerset, UK. August
Bald eagle (Haliaeetus leucocephalus) pair flying with claws linked during courtship flight, Homer, Alaska, USA, March
Portrait of a Golden snub-nosed monkey (Rhinopithecus roxellana) screaming and showing its teeth, Foping Nature Reserve, Shaanxi, China. Endangered species
Killer whale (Orcinus orca) pod hunting together in herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Playful tortoiseshell kitten
Osprey (Pandion haliaetus) in flight catching a fish, Finland, July
Wild female Amur leopard (Panthera pardus orientalis) on rocky hillside, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species
Plains zebra (Equus quagga) grooming foal, Kruger National Park, South Africa, July
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Green tree ants (Oecophylla smaragdina) defending their leaf nest, Daintree River, Wet Tropics World Heritage area, Queensland, Australia
Goldenrod crab spider (Misumena vatia) yellow female on Honesty flower, Bristol, UK, April
Japanese crane (Grus japonensis) displaying, wings in heart shape, Hokkiado, Japan, February
Great frigatebird (Fregata minor) in tree with pair of Nazca boobies (Sula granti) showing mating behaviour, Genovesa Island, Galapagos
Great spotted woodpecker (Dendrocopos major), male feeding juvenile in nest, Finland, June
Australian sea lion (Neophoca cinerea) playful juvenile swimming upside-down, Carnac Island, Western Australia
Spotted-tailed quoll (Dasyurus maculatus) scent marking in Monga National Park, New South Wales, Australia. Remote camera, triggered by movement
Common eider duck (Somateria mollissima) female defending her nest from arctic fox (Vulpes lagopus). Longyearbyen, Spitsbergen. June
Australasian crested grebe pair (Podiceps cristatus australis) in courtship display. Ashburton Lakes, Canterbury, New Zealand. August
Great crested grebe (Podiceps cristatus) performing their weed dance during courtship. Den Oever, The Netherlands. July
Veined octopus (Amphioctopus marginatus) resting on top of the two halves of an old clam shell on the sandy seabed. These octopus regularly carry pieces of shell so they can use them as protective
Murmuration of Common starlings (Sturnus vulgaris) with Peregrine falcon (Falco peregrinus) hunting them, The Netherlands, November
Mountain gorilla (Gorilla gorilla beringei) silverback Gihishamwotsi displaying, non group dominant, Sabyinyo Group, Volcanoes National Park, Virunga Mountains, Rwanda
Leopard (Panthera pardus) female fighting off male after he tries to mate with her. Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Grey whale (Eschrichtius robustus) at water surface with mouth open showing baleen plates, San Ignacio Lagoon, Baja California, Mexico
Great Bustard (Otis tarda) males displaying, Salisbury Plain, Wiltshire, UK, April
Patterns made in sand by Mint-sauce worms (Symsagittifera roscoffensis / Convoluta roscoffensis) sunbathing at low tide to allow the symbiotic algae to photosynthesise
Grizzly bear (Ursus arctos) catching a fish, Brooks Falls in Katmai National Park, Alaska, USA, July