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Horned adder (Bitis caudalis) waiting for a passing prey into the Namib Desert, Brandberg, Namibia
Horned adder (Bitis caudalis) close up of eye, Brandberg area, Namibia
Welwitschia (Welwitschia mirabilis) female plant with cones at sunset, Namib Naukluft National Park, Swakopmund, Namibia
Lioness (Panthera leo) chasing away Cheetah (Acinonyx jubatus) Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
White forest crab (Cardisoma armatum) portrait, seen through claw, Island of Principe UNESCO Biosphere Reserve, Democratic Republic of Sao Tome and Principe, Gulf of Guinea
RF - Head portrait of Spotted eagle owl (Bubo africanus), Kgalagadi Transfrontier Park, Northern Cape, South Africa, February 2016
African fish eagle (Haliaeetus vocifer). Zimanga private game reserve, KwaZulu-Natal, South Africa. June
Bateleur Eagle (Terathopius ecaudatus) female, stretching wings Kgalagadi Transfrontier Park, South Africa. January
Hawksbill turtle (Eretmochelys imbricata) feeding on soft coral (Litophyton arboreum) Nuweiba, Sinai, Egypt. Gulf of Aqaba, Red Sea
Chocolate-dip chromis (Chromis dimidiata) male over out of focus hard coral (Porites nodifera). Sha ab Claudia, Fury Shoal, Egypt. Red Sea
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon, on coral reef under stormy seas. St Johns Reef. Egypt. Red Sea
Whitetip shark (Carcharhinus longimanus), accompanied by Pilotfish (Naucrates ductor), swims towards the camera as it cruises beneath the surface of the Red Sea, close to Little Brother Island
Parachute daisies (Ursinia anthemoides) Little Karoo, Western Cape, South Africa
Massive sand dunes of Namib Naukluft National Park, Namibia
RF- Two Greater flamingos (Phoenicopterus ruber) flying across sunset sky, Namibia
Parachute daisy (Ursinia anthemoides) in flower, Vergalegen, Little Karoo, western Cape, South Africa, July
African elephant silhouetted at sunrise in the Masai Mara NR, Kenya
Kelp gull (Larus dominicanus) on rocky shore with crashing waves at sunset, Bird Island, Lamberts Bay, Western Cape province, South Africa, September 2012
Masai giraffes (Giraffa camelopardalis tippelskirchi), at sunrise, Masai-Mara Game Reserve, Kenya. March
Emperor angelfish (Pomacanthus imperator) swimming past soft corals (Dendronephthya sp]), Red Sea, Egypt. January
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at sunrise. Kasanka National Park, Zambia
Indochinese tiger (Panthera tigris corbetti) looking up, behind screen of leaves, captive, occurs in South East Asia
Eastern green mamba {Dendroaspis angusticeps} skin detail, captive, from East Africa
RF - Young lionfish (Pterois volitans) swimming near surface hunting silversides, at dusk. Beacon Rock, Sha ab Mahmood, Sinai, Egypt. Red Sea
Napoleon wrasse (Cheilinus undulatus) portrait. Red Sea, Egypt. October
Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
School of Bohar snappers (Lutjanus bohar) Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
Male Scalefin anthias (Pseudanthias squamipinnis) swimming in open water, Ras Katy, Sharm El Sheikh, Sinai, Egypt. Gulf of Aqaba, Red Sea
Lava Lake at night in the crater of Nyiragongo Volcano, Virunga National Park, North Kivu Province, Democratic Republic of Congo. September 2015
Verreauxs sifaka (Propithecus verreauxi) running across a road with bus in background, Berenty Private Reserve, southern Madagascar, August 2016
Cheetah (Acinonyx jubatus) cub aged 6 months yawning, Masai-Mara Game Reserve, Kenya. Vulnerable species
Masai giraffes (Giraffa camelopardalis tippelskirchi) in dry season, Masai-Mara Game Reserve, Kenya. March
African elephant (Loxodonta africana) herd silhouetted in distance walking through a storm, Masai-Mara Game Reserve, Kenya. September
Cheetah (Acinonyx jubatus) resting on a termite hill and storm brewing, Masai-Mara Game Reserve, Kenya. March
Cheetah (Acinonyx jubatus) female with cubs sitting in long grass during the dry season, Masai-Mara Game Reserve, Kenya. March
Wildebeest (Connochaetes taurinus) herd walking in a line in the rain at sunset, Masai-Mara Game Reserve, Kenya. September
Topi (Damaliscus korrigum / Damaliscus lunatus) territorial male standing on mound looking across savanna. Masai-Mara game reserve, Kenya
Cheetah (Acinonyx jubatus) female standing below dark storm clouds, Masai-Mara game reserve, Kenya
Springbok (Antidorcas marsupialis) grazing in front of sand dunes, Namib Naukluft Park, Namibia, June 2006
Dromedary Camel (Camelus dromedarius) on Erg Chebbi Dunes. Sahara Desert, Morocco, North Africa, March 2011
Buttress roots of rainforest tree, Loango National Park, Gabon. Photograph taken on location for BBC Africa series, January 2011
Giant groundsel (Dendrosenecio sp) at 4000m altitude on slopes of Mount Kilimanjaro, Tanzania
Massing herds of White bearded wildebeest (Connochaetes taurinus albojubatus) on migration, Serengeti National Park, Tanzania
Aerial view of Sossusvlei, Namib desert, Namibia, South Africa
Sidewinder / Peringueys adder (Bitis peringueyi) lying in wait, buried in sand ready to ambush prey, Namib Desert, Namibia
Fennec Fox (Fennecus / Vulpes zerda) in profile against sand. Dilia Achetinamou Niger, Africa