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Black-headed gull (Chroicocephalus ridibundus) in flight, artistically blurred photograph, Cheshire, UK, February. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Clarks Grebe (Aechmophorus clarkii) with yawning chick riding on back, Bear River Refuge, Utah, July
RF - Brown hare (Lepus europaeus) adult in arable field, Scotland, UK, August
Great potoo (Nyctibius grandis) female with young resting on a branch, Pantanal, Mato Grosso, Brazil
Tawny owl (Strix aluco) flying with Dormouse prey (Muscardinus avellanairus) to nest, Sussex, UK, April
Peregrine falcon (Falco peregrinus), adult male in flight. Bristol, UK. March
Goldfinch (Carduelis carduelis) adult feeding on teasel seeds, UK, February
Iberian ibex (Capra pyrenaica), adult male, walking along mountain ridge. Sierra Nevada National Park, Andalusia, Spain. June
Robin (Erithacus rubecula) singing on bare tree branch, Whitlingham Country Park, Norfolk, UK. February
Iberian ibex (Capra pyrenaica), adult male, in snow covered landscape after snowfall. Sierra Nevada National Park, Andalusia, Spain. February
Whooper swans (Cygnus cygnus) pair engaged in courtship as snow falls, Myvatn lake, Iceland, March
Digital composite of Swallowtail butterfly (Papilio machaon) caterpillar, pupa, chrysalis and emerging adult
Great horned owls (Bubo virginianus), adult and chick, on nest in Saguaro cactus (Carnegiea gigantea), Sonoran desert, Arizona
Worrells turtle (Emydura subglobosa worrelli), large adult female resting at night, Lawn Hill Creek, Queensland, Australia. January
Stag beetle (Lucanus cervus), adult male on oak tree, Italy
RF - Indian rhinoceros (Rhinoceros unicornis) Kaziranga National Park, Assam, India
RF - Brunnichs guillemot (Uria lomvia) group flying in formation, Svalbard, Norway
Polar bear (Ursus maritimus), adult male, outside Kaktovik, Alaska, USA
American Buffalo or Bison (Bison bison) group with calves, Yellowstone National Park, Wyoming, USA, May
RF- Male Emperor Dragonfly (Anax imperator) in flight, Arne RSPB reserve, Dorset, England, UK, July. Did you know? Dragonflies spend up to six years as drab aquatic nymphs for only a few weeks as
Fawn Pug dog and puppy, 8 weeks
Bengal Tiger (Panthera tigris tigris) mother and adolescent daughter fighting, Ranthambhore NP, Rajasthan, India
Red fox (Vulpes vulpes) cub biting tail of adult male, Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East. June
Galapagos giant tortoise (Chelonoidis nigra) portrait, Santa Cruz Island, Galapagos Islands, Ecuador. Endangered
Eastern rockhopper penguin (Eudyptes chrysocome filholi) mid-fall after jumping off rocky ledge to get to sea, surrounded by Bull kelp (Durvillaea antarctica), Auckland Island, Auckland Island
Florida manatees (Trichechus manatus latirostris) gathering at Three Sisters Spring, Crystal River, Florida, USA. Endangered
Great white shark (Carcharodon carcharias) swimming under the ocean surface in open water, Guadalupe Island, Mexico, Pacific Ocean
Two Iberian ibex (Capra pyrenaica), adult males, fighting. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), adult male, sitting down in snow covered landscape during snowfall. Sierra Nevada National Park, Andalusia, Spain. April
Iberian ibex (Capra pyrenaica), adult female, walking along coastal cliff. Mediterranean coast, Andalusia, Spain. November
South Andean deer (Hippocamelus bisulcus), adult female with fawn, resting on mountainside. Cerro Castillo National Park, Patagonia, Chile. April
Red-footed booby (Sula sula) perched on Palo santo tree (Bursera graveolens), with flowering Yellow cordia (Cordia lutea) in background. Galapagos Islands, Ecuador
Water rail (Rallus aquaticus) wading through water, Dumfries and Galloway, Scotland, UK
Arctic fox (Vulpes lagopus) resting in grass, Isfjorden, Svalbard, Norway, July
Walrus (Odobenus rosmarus) male resting in water, Kapp Lee, Svalbard, Norway, Arctic Ocean, July
Adult American oystercatcher (Haematopus palliatus) walking along beach, with chick running and flapping its wings. Long Island, New York, USA. June
Adult Common tern (Sterna hirundo) brooding two chicks. Long Island, New York, USA. June
Two Common terns (Sterna hirundo) in middle of aggressive encounter as one lands near nest with chick in it. Long Island, New York, USA. June
Small pearl-bordered fritillary (Boloria selene) roosting on ribwort plantain (Plantago lanceolata), Marsland mouth, Devonshire, UK. May
Small white butterfly (Pieris rapae) resting on bluebell (Hyacinthoides non-scripta), Boscastle, Cornwall, UK. April
Small tortoiseshell butterfly (Aglais urticae) basking on Cornish slate wall, Crantock, Cornwall, UK. June
Silver studded blue butterfly (Plebejus argus) resting on grass blade, Gwithian Towans, Cornwall, UK. June
Gemsbok (Oryx gazella) pair standing in grass after wet season in desert, Namib-Naukluft National Park, Namibia
Red rock crab (Grapsus adscensionis) grazing on algae from rock, Tenerife, Canary Islands, October
Brown bear (Ursus arctos) walking across rock in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. September
Sika deer (Cervus nippon) doe standing on rocky outcrop overlooking mountain forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. January
Sika deer (Cervus nippon) stag wading through river in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) stag walking on mountain slope at the edge of woodland, with forested mountains in the background, Land of the Leopard National Park, Russian Far East