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Bottle-nosed dolphins (Tursiops truncatus) breaching, Fortrose, Moray Firth, Scotland, UK, August 2010
Greater flamingos (Phoenicopterus roseus) taking off from lagoon, Camargue, France, May 2009
Two male Alpine ibex (Capra ibex ibex) fighting in front of the Mer de Glace glacier, with a female watching, Aiguilles Rouges (Red Peaks) Regional Natural Park, Haute-Savoie, France, June
Fallow deer (Dama dama) one deer stops and looks whilst the herd runs on, with sunlit leaves out of focus, Vastmanland, Sweden
Bengal Tigers (Panthera tigris) sub-adults, approximately 17-19 months old, playfighting. Endangered. Bandhavgarh National Park, India. Non-ex. Sequence 1 of 2
Lion (Panthera Leo) male running, South Africa taken on location for Pride tv series
Brown Hare (Lepus europaeus) running. Derbyshire, UK, March
Great Grey Owl (Strix nebulosa) in flight over snow covered ground, Tornio, Finland, March
Red squirrel (Sciurus vulgaris) jumping, Cairngorms National Park, Scotland, March 2012
Kingfisher (Alcedo atthis) flying up from water with caught fish, Worcestershire, UK, March
Peregrine falcon (Falco peregrinus) in flight over marshes with Golden plovers also in sky, Norfolk, UK
A bull California sealion (Zalophus californianus) bursting through a school of baitfish in the shallows. Los Isotes, La Paz, Mexico. Sea of Cortez, East Pacific Ocean
Silhouette of Red-fronted brown lemur (Eulemur rufus) leaping across gap in trees by moonlight, cathemeral behaviour, Ranomafana National Park, south east Madagascar, November
Female African lion (Panthera leo) jumping in to the Khwai river, Okavango Delta, Moremi reserve, Botswana, sequence 1/5
Wildebeest (Connochaetes taurinus) migration, herd crossing the Mara river, Masai-Mara Game Reserve, Kenya
White tailed Sea Eagle (Haliaeetus albicilla) in flight, Norway, July. Bookplate from Danny Greens The Long Journey North
Capercaillie (Tetrao urogallus) in flight, Kuusamo, Finland, January
Snow Geese (Chen caerulescens) in flight, silhouetted against colourful dusk sky. Bosque del Apache, New Mexico, USA, November
Snow Goose (Chen caerulescens), coming in to land. Bosque del Apache, New Mexico, USA, November
Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA
Little egret (Egretta garzetta) attempting to catch fish, Camargue, France, July
Great Egrets (Ardea alba) territorial fight above nest colony, Everglades National Park, Florida, USA, February
White bellied woodstar (Chaetocercus mulsant) three hovering around red flowers, Guango private reserve, Papallacta Valley, Andean Cloud Forest, East slope, Ecuador
Purple gallinule (Porphyrio martinicus) moving across waterlily covered surface, Everglades National Park, Florida, USA, March
Black faced Impala (Aepyceros melamis petersi) female jumping, Etosha National Park, Namibia
A band of semi-feral Quarter mares and foals running across a creek, Estancia Don Amerigo, Chaco, Paraguay, January 2012
Goldfinches (Carduelis carduelis) squabbling near seed feeder, Berwickshire, Scotland, April
Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Brown Long-eared bat (Plecotus auritus) drinking from a lily pond, Surrey, UK. Overall WINNER of the Audubon Society of Greater Denver Share the Viewae competition 2011
Two Scarlet macaws (Ara macao) flying, captive
A rare Sorraia stallion cantering, Coudelaria Nacional (National Stud Farm), in Alter do Chao, District of Portalegre, Alentejo, Portugal
Wild / feral Dulmen ponies (Equus caballus) close up of herd of mares and foals running on the Duke of Croys estate, Meerfelder Bruch, North Rhine-Westphalia, Germany, May 2011
Jackdaws (Corvus monedula) abstract view of flock in flight, Derbyshire, UK, December. Non-ex
Grey seal (Halichoerus grypus) adult female among breaking waves, Lincolnshire, UK Non-ex
Border Collie bitch, running towards the camera
Black Rhinoceros (Diceros bicornis) charging. Etosha National Park, Namibia, January
West Highland White Terrier leaping
Border Collie having a good shake
Red Fox (Vulpes vulpes) pouncing in grass. Vosges, France, July
Stellers Sealion (Eumetopias jubata) stampede into the water. Clayoquot Sound, Vancouver Island, Canada, March
Greylag Goose (Anser anser) landing on water. Norfolk, UK, Europe, April
Chestnut Paso peruvian stallion running, California, USA
Andalusian mare running in snow storm, Colorado, USA
Atlantic salmon (Salmo salar) moving upriver to spawn. Lligwy River nr Betws y Coed, Gwynedd, North Wales, UK, November
Brown bear (Ursus arctos) running through water, fishing, Kamchatka, Far east Russia, August
Wood Ant (Formica rufa) workers on top of their nest synchronise ejection of formic acid droplets to maximise deterrent effect on potential predator, UK
Feral domestic horses (Equus caballus) wild herd stallion (grey) fighting a bachelor chestnut stallion, Blue Swallow Reserve, near Kaapsehoop, Mpumalanga province / Eastern Transvaal, South Africa
Sri Lankan Leopard (Panthera pardus kotiya) soft focus running, captive