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Translucent coral goby (Bryaninops erythrops) in Seapen (Virgularia gustaviana). Komodo National Park, Indonesia. Indo-West Pacific
Moluccan cardinalfish (Ostorhinchus moluccensis) next to hydroids, brittlestars and sea squirts. Tulamben, Bali, Indonesia
Parrotfish (Scarus / Chlorurus sp. ) sleeping at night, New Caledonia, Pacific OceanParrotfish (Scarus / Chlorurus sp.) sleeping at night, New Caledonia, Pacific Ocean
Four Pygmy seahorses (Hippocampus bargibanti) on a Seafan / Gorgonian (Muricella sp. ) including one or several pregnant males. New Caledonia, Pacific Ocean
Scalefin anthias (Pseudanthias squamipinnis) male with a long exposure. Ras Katy, Sharm El Shiekh, Sinai, Egypt. Red Sea. Commended in the GDT Photography competition 2019
Yellow pygmy goby (Lubricogobius exiguus) portrait, Kashiwajima, Japan
Short-snouted seahorse (Hippocampus hippocampus) portrait. Tenerife, Canary Islands
Canary fish (Epinephelus marginatus) in reef. El Hierro, Canary Islands
Caribbean reef shark (Carcharhinus perezi) patrolling coral reef with Schoolmaster snapper (Lutjanus apodus) and other fish. Caribbean Sea off Gardens of the Queen National Park, Cuba
Ornamental koi carp (Cyprinus carpio) group with open mouths at lake surface. Xishuangbanna Tropical and Flowers Garden, Jinghong, Yunnan, China
Atlantic salmon (Salmo salar) on breeding territory in the River Ness, Scotland, UK, January
Big Gulf grouper (Mycteroperca jordani), Cabo Pulmo Marine National Park, Baja California Sur, Mexico
Humped rockcod (Gobionotothen gibberiformis), Antarctic Peninsula, Antarctica
Ring-tailed / Golden cardinalfish (Ostorhinchus aureus) male incubating its eggs in its mouth, Sulu sea, Philippines
Stargazer (Uranoscopus sulphureus) half buried in the sand, Sulu sea, Philippines
Henshaws snake eel (Brachysomophis henshawi) burried in the sand, Sulu sea, Philippines
Lined seahorse (Hippocampus erectus) male clinging to algae in a land locked alakaline lagoon on Eleuthera Island, Bahamas
Grayling (Thymallus thymallus) in the upper reaches of the Lena River. Baikalo-Lensky Reserve, Siberia, Russia. August 2018
Forsters hawkfish (Paracirrhites forsteri) on a coral reef. Jackson Reef, Tiran, Sinai, Egypt. Strait of Tiran, Gulf of Aqaba, Egypt
Reef manta (Mobula alfredi) female swimming close to a coral reef, while cleaner wrasse (Labroides dimidiatus), tiny by comparison, pick parasites from her lips
Three Pink anemonefish (Amphiprion perideraion) living in a purple skirted magnificent sea anemone (Heteractis magnifica) on a coral reef, below a small island
Midnight snapper (Macolor macularis) shelters in a cavern on a coral reef with white soft corals (Scleronephthya sp.) and solderfish. Vavuu Atoll, Maldives. Indian Ocean
Common seahorse (Hippocampus kuda) female wrapping her prehensile tail around piece of wood on the seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef
Whitemargin stargazer (Uranoscopus sulphureus) sets an ambush in the sand at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Reef lizardifsh (Synodus variegatus) at a cleaning station, being cleaned by Holthuis anemone shrimps (Ancylomenes holthuisi). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
Atlantic salmon (Salmon salar) under river ice. Quebec, Canada. October
Portrait of a Thorny seahorse (Hippocampus histrix) near Malapascua, Philippines
Thorny or spiny seahorse (Hippocampus histrix) portrait made off Anilao, Philippines
Shortfin lionfish (Dendrochirus brachypterus). Lembeh Strait, North Sulawesi, Indonesia
Banggai cardinalfish (Pterapogon kauderni) with a Corkscrew or Long tentacle anemone (Macrodactyla doreensis). Lembeh Strait, North Sulawesi, Indonesia
Cuckoo wrasse (Labrus mixtus) swims through a kelp forest (Laminaria sp) Porthkerris, Lizard Peninsular, Cornwall, UK August
Shoal of Piraputanga, (Brycon hilarii), close to the surface, Aquario Natural, Bonito, Mato Grosso do Sul, Brazil
Grayling (Thymallus) under Ice, Lake Baikal, Siberia, Russia
Shadowfin soldierfish (Myripristis adusta) West Papua, Indonesia
Shrimpfish / Razorfish (Aeoliscus punctatus / strigatus) have adapted to a vertical posture and swim with their head downward. Lembeh, North Sulawesi, Indonesia
Tompot blenny (Parablennius gattorugine) male peers out of its den, in leg of Swanage Pier, Swanage, Dorset, UK, English Channel, August
Brandts cormorant (Phalacrocorax penicillatus) hunts for a meal in a school of Pacific chub mackerel (Scomber japonicus), beneath an oil rig
Two male Dusky groupers (Epinephelus marginatus) fighting for territory during mating season at baixa do meros, a small pinnacle at Formigas islets Protected area, near Santa Maria Island, Azores
Shadowfin soldierfish (Myripristis adusta) schooling over coral reef. Dampier Strait, Raja Ampat, West Papua, Indonesia, March 2016
Blind loach (Nemacheilus starostini ) living in cave in the East of Turkmenistan
Wolf fish (Anarhichas lupus) portrait, Akuyeri, Iceland. North Atlantic Ocean
Arctic grayling (Thymallus arcticus) displaying his vibrant dorsal fin. North Park, Colorado, USA
False clown anemonefish (Amphiprion oceallaris) guarding their eggs laid on a coral rock beneath a sea anemonePhilippines. Bohol Sea, Tropical West Pacific Ocean
Mobula / Chilean devil ray (Mobula tarapacana) with a school of fish (Capros aper) showing vertical migration, Santa Maria, Azores, Portugal, Atlantic Ocean
Crocodilefish (Cymbacephalus beauforti) close up detail of the retractable eye-lashes, Mabul Island, Sabah, Borneo, Malaysia Celebes Sea
Soft corals (Dendronephthya hemprichi) growing in very shallow water in the shade provided by a jetty, while a school of Silversides (Atherinomorus lacunosus) circle. Berenice Jetty, Aqaba, Jordan
