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Amazon horned frog (Ceratophrys cornuta) portrait, Yasuni National Park, Orellana, Ecuador
Two paint horses, a palomino and a sorrel quarter horse running, Flitner Ranch, Shell, Wyoming, USA
Hermit thrush (Catharus guttatus) feeding on blue juniper berry (cone) of Eastern Red-Cedar (Juniperus virginiana) in mid-November, Freeville, New York, USA.
Splendid leaf frog (Cruziohyla calcarifer) La Selva Field Station, Costa Rica
Aerial view of the front of the Sermeq Kujalleq Glacier, Greenland, entering the Kangia Ilulissat Icefjord full of icebergs
Madagascan reed frog {Heterixalus madagascar- iensis} Masoala NP, Madagascar
Painted mantella frog (Mantella madagascariensis) Mantadia NP, Eastern Madagascar
Bornean horned frog (Megophrys nasuta) on forest floor. Danum Valley, Sabah, Borneo, Malaysia
Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016
Five Cavalier King Charles Spaniel sitting, black-and-tan, tricolour, blenheim and ruby coated, sitting in autumn leaves
Brown bear (Ursus arctos) and Grey wolf (Canis lupus) together in wetlands, Kuhmo, Finland, July
Duckling swimming on water surface, captive, UK
Brown Roan Italian Spinone puppy, Riley, 13 weeks, chewing his tail
Cavalier King Charles Spaniel, male with ruby coat
Red squirrel (Sciurus vulgaris), two curled up asleep in drey inside nest box. Nest of lichen and Pine needles. Highlands, Scotland, UK. Medium repro only
Polar bear (Ursus maritimus) on an iceberg, Baffin Bay, Canada. September
Pair of European grey wolves (Canis lupus) interacting, Tromso, Norway, captive, April
Cavalier King Charles Spaniel, five puppies in basket, one with tricolour and the others with blenheim coat
Gemsbok (Oryx gazella) in sand dunes, Namibia. Finalist in the Mammals Category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
Sperm whales (Physeter macrocephalus) resting, Pico, Azores, Portugal, June 2009 WWE OUTDOOR EXHIBITION. NOT AVAILABLE FOR GREETING CARDS OR CALENDARS. WWE BOOK
Shoebill stork (Balaeniceps rex) in the swamps of Mabamba, Lake Victoria, Uganda
Atlantic puffin (Fratercula arctica) landing, wings outstretched. Machias Seal Island, off the coast of Maine, USA. July
Common loon (Gavia immer) with a chick alongside. British Columbia, Canada. June
Portrait of German Shorthair Pointer in woodland, Illinois, USA
American bison (Bison bison) female covered in hoar frost near hot spring, portrait. Midway Geyser Basin, Yellowstone National Park, USA. February
Acorn Woodpecker (Melanerpes formicivorus), male at granary tree showing many acorns stored for winter survival, Orange County, California, USA
A jogger on the towpath of the Canal du Midi near Castelnaudary, Languedoc-Rousillon, France. Septeber 2011
Waterlily (Nymphaea alba) flower which has opened underwater in a lake. Alps, Ain, France, June
Herd of American bison (Bison bison) in snow, Yellowstone National Park, Wyoming, Yellowstone, January
Grizzly bear (Ursus arctos horribilis) female with cub riding on back, Katmai National Park, Alaska, USA, August
Aardvark (Orycteropus afer), young individual walking, Nambia. Captive, rescued individual
Killer whale (Orcinus orca) male named Mel attacking young South American sea lion (Otaria flavescens) on beach, Punta Norte, Peninsula Valdes, Argentina
Giant pangolin (Manis gigantea) walking along ground, Tanzania
Harpy Eagle (Harpia harpyja) portrait. Gamboa, Soberania National Park, Panama
Aerial view of river tributaries, saltmarsh and coast, Punta Umbria, Costa de la Luz, Huelva, Andalucia, Spain, March 2008
Taughannock Falls, near Ithaca, New York, in autumn after a day of heavy rainfall. October 2010. At 215 feet, Taughannock Falls is 33 feet taller than Niagara Falls
Masai giraffe {Giraffa camelopardalis} mother nuzzling baby, Lower Masai Mara GR, Kenya
Male Pileated Woodpecker (Dryocopus pileatus) with beetle larva in beak about to feed two chicks, New York, USA, May
Duckling swimming on water surface, split level, captive, UK
Underwater river scene with freshwater plants and Tetra fish, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul
Santa Cruz Galapagos tortoise (Chelonoidis nigra porteri) feeding on grass, Santa Cruz Highlands, Galapagos
Aerial view of boats and snorkelers, Looe Key Reef, Florida Keys National Marine Sanctuary, USA, showing spur and groove coral formations
Seven Ravens (Corvus corax) perched on dead tree in the snow, Utajarvi, Finland, February
Axolotl (Ambystoma mexicanum), white or leucistic form, neotenic salamander. Captive, critically endangered in the wild, occurs in Mexico
Axolotl {Siredon / Ambystoma mexicanum} albino, captive
Common opossum {Didelphis marsupialis} playing dead as defence strategy
Raccoon (Procyon lotor) peering out from behind tree trunk, Baxter State Park, Maine, USA
German shorthair pointer puppy, Pomfret, Connecticut, USA