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Alpine ibex (Capra ibex) female, Jura, Switzerland, September
Rufous motmot (Baryphthengus martii) perched on branch. Rancho Naturalista, Cartago, Costa Rica
Sea otter (Enhydra lutris) group of four resting among sea ice, Alaska, USA, June
Red deer (Cervus elaphus) stag in upland landscape. Lochcarron, Highlands, Scotland, UK
Silky shark (Carcharhinus falciformis). Cocos Island National Park, Costa Rica
Central American spider monkey (Ateles geoffroyi) juveniles hanging by tails and playing, Calakmul Biosphere Reserve, Yucatan Peninsula, Mexico, August
Sphinx hawk moth (Hemeroplanes triptolemus) caterpillar, snake mimic species, Amarakaeri Communal Reserve, Maijuna, Rio Napo, Peru
Bottlenose dolphin (Tursiops truncatus) porpoising, Sado Estuary, Portugal. October
Pine marten (Martes martes) foraging in Pine (Pinus sp) woodland at night. Glenfeshie, Cairngorms National Park, Scotland, UK
Osprey (Pandion haliaetus) fishing. Rothiemurchus, Cairngorms National Park, Scotland, UK
Ancient Scots pine (Pinus sylvestris) amongst flowering Common heather / Ling (Calluna vulgaris). Rothiemurchus Forest, Cairngorms National Park, Scotland, UK
Red fox (Vulpes vulpes) cub sitting in coniferous woodland clearing at dusk. Glenfeshie, Cairngorms National Park, Scotland, UK
Bigeye trevally (Caranx sexfasciatus), a large shoal. Cocos Island National Park, Costa Rica
Eurasian badger (Meles meles), two foraging in Pine (Pinus sp) woodland. Glenfeshie, Cairngorms National Park, Scotland, UK
Rocks and coastal rainforest. Cocos Island National Park, Costa Rica. 2018
Marine iguana (Amblyrhynchus cristatus) basking on volcanic rock at coast. Isabela Island, Galapagos
Blue-lined snapper (Lutjanus kasmira) shoal. Cocos Island National Park, Costa Rica
Fish seeking shelter around floating tree in open ocean. Off coast of Cocos Island National Park, Costa Rica
Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica
Rivolis hummingbird (Eugenes fulgens) nectaring on Fuchsia (Fuchsia sp) flower. Costa Rica
Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica
Violet sabrewing hummingbird (Campylopterus hemileucurus) nectaring on Orchid. Costa Rica
Devils hand Tree (Chiranthodendron pentadactylon) flower, Filo de Caballos, Sierra Madre del Sur, southern Mexico, March
Sugar kelp (Saccharina latissima) and Sea anemones (Actiniaria) on rock near iceberg. Tasiilaq, East Greenland. April
Broadfoot mushroomtongue salamander (Bolitoglossa platydactyla), Catemaco Lake, Los Tuxtlas Rainforest, Mexico, July
Himalayan marmot (Marmota himalayana). Chantang Wlidlife Sanctuary. Ladakh, India
St Michaels Mount and old causeway at sunrise, Marazion, Cornwall, UK. October 2015
Lotus (Nelumbo nucifera) in flower in botanic garden, Vendee, France, July
Squirrelfish (Holocentrus sp) shoal amongst rocks. Near Cocos Island National Park, Costa Rica
Galapagos green turtle (Chelonia mydas agassizi) swimming above sea floor with shoal of fish in background. Off San Cristobal Island, Galapagos
Galapagos green turtle (Chelonia mydas agassizi) swimming in coastal waters of San Cristobal Island, Galapagos
Sergeant major (Abudefduf sp) schooling in mangroves of Isabela Island, Galapagos
Red squirrel (Sciurus vulgaris) feeding in snow. Scotland, UK. March
Red squirrel (Sciurus vulgaris) jumping towards camera. Scotland, UK. February
Red deer (Cervus elaphus) herd during rut in morning light. Derbyshire, England, UK. October
Resplendent quetzal (Pharomachrus mocinno) male perched on branch. Costa Rica
Marsh fritillary (Euphydryas aurinia), group feeding on Marsh thistle (Cirsium palustre). Lincolnshire, England, UK
Leaf-nosed bat (Phyllostomidae sp) flying towards Banana (Musa sp) flower to feed. Costa Rica
Green-crowned brilliant (Heliodoxa jacula), three feeding on Lobster-claw (Heliconia sp) flower in rain. Costa Rica
Red fox (Vulpes vulpes), two cubs play fighting. Sheffield, England, UK. May
Red fox (Vulpes vulpes) cub looking over shoulder at camera. Sheffield, England, UK. May
Red fox (Vulpes vulpes) cub looking at camera, in morning. Sheffield, England, UK. June
Badger (Meles meles) eating apples in urban garden. Sheffield, England, UK. October
Badger (Meles meles) amongst Bluebells (Hyacinthoides non-scripta). Scotland, UK. May
Badger (Meles meles) emerging through gateway in wall at night. In urban area, Sheffield, England, UK. October 2018
Wood warbler (Phylloscopus sibilatrix) singing whilst perched on branch, silhouetted at dawn. Sheffield, England, UK. May
Turtle dove (Streptopelia turtur) standing at a drinking pool. Hungary. May
Spoonbill (Platalea leucorodia) preening, close up. Hungary. May
Aerial view of wetlands landscape, Wild Grey wolf (Canis lupus) habitat, Saxony, Germany, September
Red deer (Cervus elaphus) stag, Saxony, Germany. September
Wild Grey wolf (Canis lupus), in meadow, Saxony-Anhalt, Germany
Wild Grey wolf (Canis lupus) walking, Saxony-Anhalt, Germany
Mustached Bat (Pteronotus sp. ) flying at duskMustached Bat (Pteronotus sp.) flying at dusk, Calakmul Biosphere Reserve, Yucatan Peninsula, Mexico, August
Puente de Dios cascades, Tamasopo, Huasteca Region, Mexico, August
Sugar kelp (Saccharina latissima) near iceberg, view upwards to gap in sea ice. Tasiilaq, East Greenland. April
Female House spider (Tegenaria sp)Female House spider (Tegenaria sp.) at the mouth of her tubular silk retreat in an old stone wall, Somerset, UK, September
Black-headed gull (Chroicocephalus ridibundus) flock flying over a frozen lake in falling snow, Wiltshire, UK, March
Running foliage spider / Spiny-legged sac spider (Liocranum rupicola), a nationally scarce species in the UK, hunting at night in an old stone wall, Wiltshire, UK, September
Common lace weaver / Lace-webbed spider (Amaurobius similis) male in its web in an old stone wall at night, Wiltshire, UK, September
Hornet (Vespa crabro) drinking at bird bath. Dorset, UK, July
Red fox (Vulpes vulpes) cubs playing, Vosges, France, May
Wild cat (Felis silvestris) walking, Vosges, France, June
Jewel beetle (Anthaxia hungarica) on aster flower, South of Casteil, French Pyrenees, France. May
Red fox (Vulpes vulpes) cub, Vosges, France, May
Herald moth (Scoliopteryx libatrix) camouflaged in leaf litter, Gosford Forest Park, Co. Armagh, Northern Ireland. September
Spotted eagle owl (Bubo africanus) subadult at nigh by water, Zimanga private game reserve, KwaZulu-Natal, South Africa, September
Great tiger moth (Arctia caja americana) Lac-Drolet, province, Quebec, Canada November
Leopard / mountain tortoise (Stigmochelys pardalis) with elephant bull (Loxodonta africana) behind in water, Addo elephant national park, Eastern Cape, South Africa, September
Impala (Aepyceros melampus) allo-grooming, iMfolozi game reserve, KwaZulu-Natal, South Africa, August
Giraffe (Giraffa camelopardalis) drinking at night, Zimanga private game reserve, KwaZulu-Natal, South Africa August
Malachite sunbird (Nectarinia famosa) feding at flower, Kirstenbosch National Botanical Garden, Cape Town, South Africa, September
Kiangs (Equus kiang) on grassland near Tso Kar Lake, Chantang Wildlife Sanctuary. Ladakh, India, September 2018
Orange-breasted sunbird (Anthobaphes violacea) female on Leucospermum cordifolium Yellow Bird Kirstenbosch National Botanical Garden, Cape Town, South Africa, September
Cape sugarbird (Promerops cafer) on king protea, Kirstenbosch National Botanical Garden, Cape Town, South Africa, September
Southern double-collared sunbird (Cinnyris chalybeus), Kirstenbosch National Botanical Garden, Cape Town, South Africa, September
Rainbow trout (Oncorhynchus mykiss) in green turbid waters, Gunnison River, Colorado, USA, April
Bactrian camels (Camelus bactrianus) on sand, Nubra Valley, Ladakh, India. September
Holywell Bay, low tide at sunset, Carters Rocks and reflections, near Newquay, Cornwall, UK. April 2017
The Old Man of Storr, golden early morning light, the Trotternish peninsula, Isle of Skye, Scotland, UK. November 2017
The Quiraing in morning light, eastern face of Meall na Suiramach, the northernmost summit of the Trotternish on the Isle of Skye, Scotland, UK. November 2017
Black-legged kittiwake (Rissa tridactyla) in flight over Newcastle city centre in a blizzard. Newcastle, UK. March
Red-legged partridge (Alectoris rufa) with chicks, Vendee, France, June
Red fox (Vulpes vulpes) and a newborn cub, Sado Estuary, Portugal. May